About
Education
Discuss
TraitBank
Sign In
Sign Up
Language
Deutsch
English
Español
français
italiano
Nederlands
Piemontèis
Português do Brasil
suomi
Türkçe
čeština
Ελληνικά
македонски
Українська
العربية
简体中文
繁體中文
names in breadcrumbs
vernacular
scientific
About
Education
Discuss
TraitBank
Sign In
Sign Up
en
Deutsch
English
Español
français
italiano
Nederlands
Piemontèis
Português do Brasil
suomi
Türkçe
čeština
Ελληνικά
македонски
Українська
العربية
简体中文
繁體中文
names in breadcrumbs
vernacular
scientific
Creatures
»
…
»
Animal
»
…
»
Arthropods
»
…
»
Hexapods
»
Insects
»
Winged Insects
»
…
»
Endopterygotes
»
Beetles
»
…
»
Scarab Beetles
Creatures
»
Cellular Organisms
»
Eukaryotes
»
Opisthokonts
»
Animal
»
Bilateria
»
Protostomes
»
Ecdysozoans
»
Arthropods
»
Pancrustacea
»
Hexapods
»
Insects
»
Winged Insects
»
Neopteran
»
Endopterygotes
»
Beetles
»
Polyphaga
»
Scarabaeiformia
»
Scarab Beetle
»
Scarab Beetles
«
Parascatonomus
collect
overview
data
media
articles
maps
names
Scientific Names
Preferred Names
Parascatonomus
Genus recognized by
EOL Dynamic Hierarchy April 2022
Reference taxon from Catalogue of Life in Species 2000 & ITIS Catalogue of Life
Parascatonomus
Genus recognized by
EOL Dynamic Hierarchy 1.1
,
EOL Dynamic Hierarchy 1.1
,
Vietnam Species List
,
Japan Species List
,
North Pacific Species List
,
NCBI
,
wikidata_hierarchy tar gz
,
iNaturalist data coverage
,
Dynamic Hierarchy Version 2.2.3
, and
EOL Dynamic Hierarchy Version 2.2.4
Parascatonomus
Subgenus recognized by
wikipedia NO
and
wikidata_hierarchy tar gz
Parascatonomus Paulian 1932
Genus recognized by
GBIF classification
,
Catalogue of Life 26 March 2024
, and
GBIF data coverage
Parascatonomus subgen. Parascatonomus Paulian 1932
Subgenus recognized by
Catalogue of Life 26 March 2024
Alternative Names
Neosisyphus penicillatus
(Harold 1880)
Homotypic synonym according to
GBIF classification
(Originally found in sources as accepted taxon within Neosisyphus Müller, 1942)
Onthophagus acuticollis
Gillet
Homotypic synonym according to
GBIF classification
(Originally found in sources as accepted taxon within Onthophagus Latreille, 1802)
Onthophagus anguicorius
Boucomont 1914
Homotypic synonym according to
GBIF classification
(Originally found in sources as accepted taxon within Onthophagus Latreille, 1802)
Onthophagus crassus
Sharp 1875
Heterotypic synonym according to
GBIF classification
Onthophagus diversiformis
Boucomont 1914
Homotypic synonym according to
GBIF classification
(Originally found in sources as misapplied of Onthophagus chebaicus Boucomont, 1919)
Onthophagus itiokai
Ochi, Kon & Kishimoto-Yamada 2010
Homotypic synonym according to
GBIF classification
(Originally found in sources as doubtful taxon within Onthophagus Latreille, 1802)
Onthophagus kiuchianus
Masumoto, Yang & Ochi 2004
Homotypic synonym according to
GBIF classification
(Originally found in sources as accepted taxon within Parascatonomus Paulian, 1932)
Onthophagus kuatunensis
Balthasar
Homotypic synonym according to
GBIF classification
(Originally found in sources as accepted taxon within Onthophagus Latreille, 1802)
Onthophagus tricornis
Wiedemann
Homotypic synonym according to
GBIF classification
(Originally found in sources as accepted taxon within Parascatonomus Paulian, 1932)
? penicillatus Harold, 1880
Homotypic synonym according to
GBIF classification
(Originally found in sources as doubtful taxon within Scarabaeidae)
Coomanius Paulian 1932
Synonym according to
Catalogue of Life 26 March 2024
Parascatonomus
Harvest ancestor according to
iNaturalist
and
Global Biotic Interactions
Common Names
There are no common names associated with this taxon.
Curated hierarchies for Parascatonomus
EOL Dynamic Hierarchy April 2022
Life
Cellular Organisms
Eukaryota
Opisthokonta
Metazoa
Bilateria
Protostomia
Ecdysozoa
Arthropoda
Pancrustacea
Hexapoda
Insecta
Pterygota
Neoptera
Endopterygota
Coleoptera
Polyphaga
Scarabaeiformia
Scarabaeoidea
Scarabaeidae
Parascatonomus
(this page)
Parascatonomus (Granulidorsum) aper (Sharp 1875)
Parascatonomus (Granulidorsum) dejongi (Krikken 1986)
Parascatonomus (Granulidorsum) hayashimasaoi (Ochi & Kon 1999)
Parascatonomus (Granulidorsum) koichii Ochi & Araya 1992
Parascatonomus (Granulidorsum) leytensis Ochi & Araya 1992
Parascatonomus (Granulidorsum) negrosensis Ochi & Araya 1992
Parascatonomus (Granulidorsum) pseudorudis Ochi & Araya 1992
Parascatonomus (Granulidorsum) quezonensis Ochi & Araya 1992
Parascatonomus (Granulidorsum) rudis (Sharp 1875)
Parascatonomus (Granulidorsum) sakamakii (Ochi, Kon & Kawahara 2010)
Parascatonomus (Granulidorsum) shizumui Ochi & Araya 1992
Parascatonomus (Granulidorsum) takedai Ochi & Araya 1992
Parascatonomus (Granulidorsum) takeshii Ochi & Araya 1992
Parascatonomus (Granulidorsum) yvescambeforti Ochi & Araya 1992
Parascatonomus (Necramator) accedens (Lansberge 1883)
Parascatonomus (Necramator) acuticollis (Gillet 1927)
Parascatonomus (Necramator) aerumnosus (Balthasar 1963)
Parascatonomus (Necramator) alexeevi (Tarasov, Krikken & Huijbregts 2010)
Parascatonomus (Necramator) anguicorius (Boucomont 1914)
Parascatonomus (Necramator) anitidus (Ochi & Kon 2005)
Parascatonomus (Necramator) aokii (Nomura 1976)
Parascatonomus (Necramator) atratus (Kabakov 1983)
Parascatonomus (Necramator) aurifex (Harold 1877)
Parascatonomus (Necramator) brendelli Ochi, Kon & Barclay 2008
Parascatonomus (Necramator) bundutuhanensis Ochi, Kon & Barclay 2008
Parascatonomus (Necramator) chaiyaphumensis Masumoto & Ochi 2014
Parascatonomus (Necramator) chapaensis (Kabakov 1983)
Parascatonomus (Necramator) chiangraiensis (Masumoto, Ochi & Hanboonsong 2007)
Parascatonomus (Necramator) damaki (Masumoto, Hanboonsong & Ochi 2002)
Parascatonomus (Necramator) danumcupreus (Krikken & Huijbregts 2009)
Parascatonomus (Necramator) diversiformis (Boucomont 1914)
Parascatonomus (Necramator) dux (Sharp 1875)
Parascatonomus (Necramator) formosanus (Gillet 1924)
Parascatonomus (Necramator) funebris (Boucomont 1919)
Parascatonomus (Necramator) gunsalami (Ochi & Kon 2005)
Parascatonomus (Necramator) horni (Balthasar 1935)
Parascatonomus (Necramator) igneus (Vigors 1825)
Parascatonomus (Necramator) itiokai (Ochi, Kon & Kishimoto-Yamada 2010)
Parascatonomus (Necramator) itoi (Nomura 1976)
Parascatonomus (Necramator) javacupreus Krikken & Huijbregts 2009
Parascatonomus (Necramator) javanitidus (Ochi, Kon & Hartini 2010)
Parascatonomus (Necramator) katoi (Ochi & Araya 1996)
Parascatonomus (Necramator) kikutai (Ochi & Kon 2005)
Parascatonomus (Necramator) kiuchianus (Masumoto, Yang & Ochi 2004)
Parascatonomus (Necramator) klapperichi (Balthasar 1952)
Parascatonomus (Necramator) kuatunensis (Balthasar 1942)
Parascatonomus (Necramator) laevicollis (Lansberge 1883)
Parascatonomus (Necramator) lakyim (Masumoto 1990)
Parascatonomus (Necramator) liewi (Ochi & Kon 2005)
Parascatonomus (Necramator) maesaensis Masumoto & Ochi 2014
Parascatonomus (Necramator) mairuu (Masumoto 1989)
Parascatonomus (Necramator) masahiroi (Masumoto, Ochi & Hanboonsong 2008)
Parascatonomus (Necramator) monticupreus (Krikken & Huijbregts 2009)
Parascatonomus (Necramator) munetoshii Masumoto, Ochi & Sakchoowong 2012
Parascatonomus (Necramator) murasakianus (Nomura 1976)
Parascatonomus (Necramator) muticifrons (Endrödi 1973)
Parascatonomus (Necramator) naviculifer (Boucomont 1919)
Parascatonomus (Necramator) niasensis (Boucomont 1914)
Parascatonomus (Necramator) nitidus (Waterhouse 1875)
Parascatonomus (Necramator) nusatenggaricus (Krikken & Huijbreghts 2008)
Parascatonomus (Necramator) peninsulocupreus Krikken & Huijbregts 2009
Parascatonomus (Necramator) phraoensis (Masumoto 1990)
Parascatonomus (Necramator) pilularius (Lansberge 1883)
Parascatonomus (Necramator) piyawati Masumoto, Ochi & Hanboonsong 2002
Parascatonomus (Necramator) poringensis (Ochi & Kon 2005)
Parascatonomus (Necramator) quaestus (Sharp 1875)
Parascatonomus (Necramator) rhombocephalus (Kabakov 1983)
Parascatonomus (Necramator) salakensis (Ochi, Kon & Hartini 2010)
Parascatonomus (Necramator) sarawacus (Harold 1877)
Parascatonomus (Necramator) satoi (Ochi, Shimada & Kon 2003)
Parascatonomus (Necramator) sayapensis (Ochi & Kon 2005)
Parascatonomus (Necramator) semiaureus (Lansberge 1883)
Parascatonomus (Necramator) semicupreus (Harold 1877)
Parascatonomus (Necramator) semifex (Krikken & Huijbregts 2008)
Parascatonomus (Necramator) serapiensis (Ochi, Kon & Barclay 2008)
Parascatonomus (Necramator) shibatai (Nakane 1960)
Parascatonomus (Necramator) striaticollis (Kabakov 1983)
Parascatonomus (Necramator) sumawacus (Krikken & Huijbregts 2008)
Parascatonomus (Necramator) taichii (Ochi, Kon & Barclay 2008)
Parascatonomus (Necramator) takahashii Masumoto, Ochi & Hanboonsong 2013
Parascatonomus (Necramator) takakui (Ochi, Kon & Hartini 2009)
Parascatonomus (Necramator) tamijii (Kon, Sakai & Ochi 2000)
Parascatonomus (Necramator) taominorui Masumoto & Ochi 2014
Parascatonomus (Necramator) thaitai (Masumoto, Ochi & Hanboonsong 2008)
Parascatonomus (Necramator) tricornis (Wiedemann 1823)
Parascatonomus (Necramator) trungboensis (Kabakov 1983)
Parascatonomus (Necramator) utsunomiyae (Masumoto, Ochi & Hanboonsong 2007)
Parascatonomus (Necramator) vasiljevi Kabakov 1992
Parascatonomus (Necramator) vitalisi (Boucomont 1921)
Parascatonomus (Necramator) vividus (Arrow 1907)
Parascatonomus (Peculiaricerus) anceyi (Boucomont 1921)
Parascatonomus (Peculiaricerus) egregius (Arrow 1907)
Parascatonomus (Pseudonthophagus) penicillatus (Harold 1879)
Parascatonomus (Pseudonthophagus) yukae (Masumoto, Ochi & Hanboonsong 2002)
Parascatonomus abramovi (Zinchenko 2018)
Parascatonomus apilularius (Masumoto 1995)
Parascatonomus balikpapanensis Ochi & Kon 2016
Parascatonomus discedens (Sharp 1875)
Parascatonomus fujiokai (Ochi & Araya 1996)
Parascatonomus gosoli (Masumoto 1989)
Parascatonomus impar (Kabakov 2006)
Parascatonomus javapolitus Ochi, Kon & Barclay 2017
Parascatonomus laotianus (Boucomont 1919)
Parascatonomus maculatus (Fabricius 1801)
Parascatonomus miyakei Ochi & Araya 1992
Parascatonomus nilgiripolitus Ochi, Kon & Barclay 2017
Parascatonomus paotao Ochi, Kon & Barclay 2017
Parascatonomus paramairuu Ochi & Kon 2016
Parascatonomus politus (Paulian 1932)
Parascatonomus seminiasensis Ochi & Kon 2016
Parascatonomus shanicus Kabakov 2006
Parascatonomus sikkimpolitus Ochi, Kon & Barclay 2017
Parascatonomus vietnampolitus Ochi, Kon & Barclay 2017
Acanthonitis
Aliuscanthoniola
Allonitis
Anoctus
Anomiopsoides
Anonychonitis
Anoplodrepanus
Apotolamprus
Attavicinus
Ausmontins
Bdelyrus
Besourenga
Byrrhidium
Cambefortantus
Cambefortius
Canthon
Canthonidia
Catharsiocopris
Catharsius
Chalcocopris
Cheironitis
Clypeodrepanus
Copridaspidus
Copris
Coprophanaeus
Delopleurus
Deltochilum
Deltorhinum
Dendropaemon
Digitonthophagus
Diorygopyx
Dorbignyolus
Drepanocerus
Drepanoplatynus
Drogo
Dwesasilvasedis
Epidrepanus
Escarabaeus
Eudinopus
Euoniticellus
Eusaproecius
Frankenbergerius
Gilletellus
Glyphoderus
Gymnopleurus
Helictopleurus
Heliocopris
Heteroclitopus
Heteronitis
Holocephalus
Homocopris
Homocorpis
Hypocanthidium
Isocopris
Jossonthophagus
Latodrepanus
Liatongus
Martinezidium
Megathopa
Melanocanthon
Mentophilus
Microcopris
Mimonthophagus
Mnematidium
Monoplistes
Morettius
Neosaproecius
Ochicanthon
Oniticellus
Onitis
Onthophagus
Oxysternon
Pachylomera
Paracanthon
Paracryptocanthon
Paronthobium
Pedaria
Phaedotrogus
Pinacopodius
Pinacotarsus
Platyonitis
Pleuronitis
Pseudocopris
Pseudonthobium
Pseudopedaria
Pseudosaproecius
Scaptocnemis
Scarabaeus
Scatonomus
Scatrichus
Sceliages
Scybalocanthon
Silvinha
Streblopus
Sulcophanaeus
Temnoplectron
Theotimius
Trichillidium
Trichillum
Tropidonitis
178 additional siblings truncated for brevity.
See the
resource file
for a full list.
Catalogue of Life 26 March 2024
Biota
Animalia
Arthropoda
Insecta
Coleoptera
Scarabaeoidea
Scarabaeidae Latreille 1802
Scarabaeinae Latreille 1802
Onthophagini Streubel 1846
Parascatonomus Paulian 1932
(this page)
Parascatonomus subgen. Granulidorsum Ochi & Kon 2017
Parascatonomus subgen. Laonthophagus Frey 1971
Parascatonomus subgen. Necramator Ochi & Kon 2017
Parascatonomus subgen. Parascatonomus Paulian 1932
Parascatonomus subgen. Peculiaricerus Ochi & Kon 2017
Parascatonomus subgen. Pseudonthophagus Balthasar 1959
Alloscelus Boucomont 1923
Amietina Cambefort 1981
Anoctus Sharp 1875
Caccobiomorphus Balthasar 1964
Caccobius Thomson 1859
Cambefortius Branco 1989
Cleptocaccobius Cambefort 1984
Cyobius Sharp 1875
Diastellopalpus Lansberge 1886
Digitonthophagus Balthasar 1959
Disphysema Harold 1873
Dorbignyolus Branco 1989
Euonthophagus Balthasar 1959
Eusaproecius Branco 1989
Hamonthophagus Roggero, Dierkens, Barbero & Palestrini 2017
Heteroclitopus Péringuey 1901
Hyalonthophagus Palestrini & Giacone 1988
Jossonthophagus Dierkens, Moretto & Génier 2017
Krikkenius Branco 1991
Kurtops Roggero, Barbero & Palestrini 2017
Larhodius Balthasar 1963
Milichus Péringuey 1901
Mimonthophagus Balthasar 1963
Morettius Roggero, Dierkens, Barbero & Palestrini 2017
Neosaproecius Branco 1990
Onthophagus Latreille 1802
Phaedotrogus Paulian 1985
Phalops Erichson 1848
Pinacopodius Branco 1991
Pinacotarsus Harold 1875
Proagoderus Lansberge 1883
Pseudosaproecius Balthasar 1941
Stiptocnemis Branco 1989
Stiptopodius Harold 1871
Stiptotarsus Branco 1989
Sukelus Branco 1992
Tiaronthophagus Roggero, Moretto, Barbero & Palestrini 2019
Tomogonus Orbigny 1904
Uenoeus Ochi & Kon 2017
Unidentis Josso & Prévost 2008
Walterantus Cambefort 1977
Catalogue of Life 26 March 2024
Biota
Animalia
Arthropoda
Insecta
Coleoptera
Scarabaeoidea
Scarabaeidae Latreille 1802
Scarabaeinae Latreille 1802
Onthophagini Streubel 1846
Parascatonomus Paulian 1932
Parascatonomus subgen. Parascatonomus Paulian 1932
(this page)
Parascatonomus (Parascatonomus) abramovi (Zinchenko 2018)
Parascatonomus (Parascatonomus) apilularius (Masumoto 1995)
Parascatonomus (Parascatonomus) balikpapanensis Ochi & Kon 2016
Parascatonomus (Parascatonomus) discedens (Sharp 1875)
Parascatonomus (Parascatonomus) fujiokai (Ochi & Araya 1996)
Parascatonomus (Parascatonomus) gosoli (Masumoto 1989)
Parascatonomus (Parascatonomus) graphicus (Wallengren 1881)
Parascatonomus (Parascatonomus) impar (Kabakov 2006)
Parascatonomus (Parascatonomus) javapolitus Ochi, Kon & Barclay 2017
Parascatonomus (Parascatonomus) laotianus (Boucomont 1919)
Parascatonomus (Parascatonomus) maculatus (Fabricius 1801)
Parascatonomus (Parascatonomus) miyakei (Ochi & Araya 1992)
Parascatonomus (Parascatonomus) nilgiripolitus Ochi, Kon & Barclay 2017
Parascatonomus (Parascatonomus) paotao Ochi, Kon & Barclay 2017
Parascatonomus (Parascatonomus) paramairuu Ochi & Kon 2016
Parascatonomus (Parascatonomus) politus (Paulian 1932)
Parascatonomus (Parascatonomus) seminiasensis Ochi & Kon 2016
Parascatonomus (Parascatonomus) shanicus Kabakov 2006
Parascatonomus (Parascatonomus) sikkimpolitus Ochi, Kon & Barclay 2017
Parascatonomus (Parascatonomus) simpliciceps (Orbigny 1905)
Parascatonomus (Parascatonomus) vietnampolitus Ochi, Kon & Barclay 2017
Parascatonomus subgen. Granulidorsum Ochi & Kon 2017
Parascatonomus subgen. Laonthophagus Frey 1971
Parascatonomus subgen. Necramator Ochi & Kon 2017
Parascatonomus subgen. Peculiaricerus Ochi & Kon 2017
Parascatonomus subgen. Pseudonthophagus Balthasar 1959
Dynamic Hierarchy Version 2.2.3
Parascatonomus
(this page)
Parascatonomus (Granulidorsum) aper (Sharp 1875)
Parascatonomus (Granulidorsum) dejongi (Krikken 1986)
Parascatonomus (Granulidorsum) hayashimasaoi (Ochi & Kon 1999)
Parascatonomus (Granulidorsum) koichii Ochi & Araya 1992
Parascatonomus (Granulidorsum) leytensis Ochi & Araya 1992
Parascatonomus (Granulidorsum) negrosensis Ochi & Araya 1992
Parascatonomus (Granulidorsum) pseudorudis Ochi & Araya 1992
Parascatonomus (Granulidorsum) quezonensis Ochi & Araya 1992
Parascatonomus (Granulidorsum) rudis (Sharp 1875)
Parascatonomus (Granulidorsum) sakamakii (Ochi, Kon & Kawahara 2010)
Parascatonomus (Granulidorsum) shizumui Ochi & Araya 1992
Parascatonomus (Granulidorsum) takedai Ochi & Araya 1992
Parascatonomus (Granulidorsum) takeshii Ochi & Araya 1992
Parascatonomus (Granulidorsum) yvescambeforti Ochi & Araya 1992
Parascatonomus (Necramator) accedens (Lansberge 1883)
Parascatonomus (Necramator) acuticollis (Gillet 1927)
Parascatonomus (Necramator) aerumnosus (Balthasar 1963)
Parascatonomus (Necramator) alexeevi (Tarasov, Krikken & Huijbregts 2010)
Parascatonomus (Necramator) anguicorius (Boucomont 1914)
Parascatonomus (Necramator) anitidus (Ochi & Kon 2005)
Parascatonomus (Necramator) aokii (Nomura 1976)
Parascatonomus (Necramator) atratus (Kabakov 1983)
Parascatonomus (Necramator) aurifex (Harold 1877)
Parascatonomus (Necramator) brendelli Ochi, Kon & Barclay 2008
Parascatonomus (Necramator) bundutuhanensis Ochi, Kon & Barclay 2008
Parascatonomus (Necramator) chaiyaphumensis Masumoto & Ochi 2014
Parascatonomus (Necramator) chapaensis (Kabakov 1983)
Parascatonomus (Necramator) chiangraiensis (Masumoto, Ochi & Hanboonsong 2007)
Parascatonomus (Necramator) damaki (Masumoto, Hanboonsong & Ochi 2002)
Parascatonomus (Necramator) danumcupreus (Krikken & Huijbregts 2009)
Parascatonomus (Necramator) diversiformis (Boucomont 1914)
Parascatonomus (Necramator) dux (Sharp 1875)
Parascatonomus (Necramator) formosanus (Gillet 1924)
Parascatonomus (Necramator) funebris (Boucomont 1919)
Parascatonomus (Necramator) gunsalami (Ochi & Kon 2005)
Parascatonomus (Necramator) horni (Balthasar 1935)
Parascatonomus (Necramator) igneus (Vigors 1825)
Parascatonomus (Necramator) itiokai (Ochi, Kon & Kishimoto-Yamada 2010)
Parascatonomus (Necramator) itoi (Nomura 1976)
Parascatonomus (Necramator) javacupreus Krikken & Huijbregts 2009
Parascatonomus (Necramator) javanitidus (Ochi, Kon & Hartini 2010)
Parascatonomus (Necramator) katoi (Ochi & Araya 1996)
Parascatonomus (Necramator) kikutai (Ochi & Kon 2005)
Parascatonomus (Necramator) kiuchianus (Masumoto, Yang & Ochi 2004)
Parascatonomus (Necramator) klapperichi (Balthasar 1952)
Parascatonomus (Necramator) kuatunensis (Balthasar 1942)
Parascatonomus (Necramator) laevicollis (Lansberge 1883)
Parascatonomus (Necramator) lakyim (Masumoto 1990)
Parascatonomus (Necramator) liewi (Ochi & Kon 2005)
Parascatonomus (Necramator) maesaensis Masumoto & Ochi 2014
Parascatonomus (Necramator) mairuu (Masumoto 1989)
Parascatonomus (Necramator) masahiroi (Masumoto, Ochi & Hanboonsong 2008)
Parascatonomus (Necramator) monticupreus (Krikken & Huijbregts 2009)
Parascatonomus (Necramator) munetoshii Masumoto, Ochi & Sakchoowong 2012
Parascatonomus (Necramator) murasakianus (Nomura 1976)
Parascatonomus (Necramator) muticifrons (Endrödi 1973)
Parascatonomus (Necramator) naviculifer (Boucomont 1919)
Parascatonomus (Necramator) niasensis (Boucomont 1914)
Parascatonomus (Necramator) nitidus (Waterhouse 1875)
Parascatonomus (Necramator) nusatenggaricus (Krikken & Huijbreghts 2008)
Parascatonomus (Necramator) peninsulocupreus Krikken & Huijbregts 2009
Parascatonomus (Necramator) phraoensis (Masumoto 1990)
Parascatonomus (Necramator) pilularius (Lansberge 1883)
Parascatonomus (Necramator) piyawati Masumoto, Ochi & Hanboonsong 2002
Parascatonomus (Necramator) poringensis (Ochi & Kon 2005)
Parascatonomus (Necramator) quaestus (Sharp 1875)
Parascatonomus (Necramator) rhombocephalus (Kabakov 1983)
Parascatonomus (Necramator) salakensis (Ochi, Kon & Hartini 2010)
Parascatonomus (Necramator) sarawacus (Harold 1877)
Parascatonomus (Necramator) satoi (Ochi, Shimada & Kon 2003)
Parascatonomus (Necramator) sayapensis (Ochi & Kon 2005)
Parascatonomus (Necramator) semiaureus (Lansberge 1883)
Parascatonomus (Necramator) semicupreus (Harold 1877)
Parascatonomus (Necramator) semifex (Krikken & Huijbregts 2008)
Parascatonomus (Necramator) serapiensis (Ochi, Kon & Barclay 2008)
Parascatonomus (Necramator) shibatai (Nakane 1960)
Parascatonomus (Necramator) striaticollis (Kabakov 1983)
Parascatonomus (Necramator) sumawacus (Krikken & Huijbregts 2008)
Parascatonomus (Necramator) taichii (Ochi, Kon & Barclay 2008)
Parascatonomus (Necramator) takahashii Masumoto, Ochi & Hanboonsong 2013
Parascatonomus (Necramator) takakui (Ochi, Kon & Hartini 2009)
Parascatonomus (Necramator) tamijii (Kon, Sakai & Ochi 2000)
Parascatonomus (Necramator) taominorui Masumoto & Ochi 2014
Parascatonomus (Necramator) thaitai (Masumoto, Ochi & Hanboonsong 2008)
Parascatonomus (Necramator) tricornis (Wiedemann 1823)
Parascatonomus (Necramator) trungboensis (Kabakov 1983)
Parascatonomus (Necramator) utsunomiyae (Masumoto, Ochi & Hanboonsong 2007)
Parascatonomus (Necramator) vasiljevi Kabakov 1992
Parascatonomus (Necramator) vitalisi (Boucomont 1921)
Parascatonomus (Necramator) vividus (Arrow 1907)
Parascatonomus (Peculiaricerus) anceyi (Boucomont 1921)
Parascatonomus (Peculiaricerus) egregius (Arrow 1907)
Parascatonomus (Pseudonthophagus) penicillatus (Harold 1879)
Parascatonomus (Pseudonthophagus) yukae (Masumoto, Ochi & Hanboonsong 2002)
Parascatonomus abramovi (Zinchenko 2018)
Parascatonomus apilularius (Masumoto 1995)
Parascatonomus balikpapanensis Ochi & Kon 2016
Parascatonomus discedens (Sharp 1875)
Parascatonomus fujiokai (Ochi & Araya 1996)
Parascatonomus gosoli (Masumoto 1989)
Parascatonomus impar (Kabakov 2006)
Parascatonomus javapolitus Ochi, Kon & Barclay 2017
Parascatonomus laotianus (Boucomont 1919)
Parascatonomus maculatus (Fabricius 1801)
Parascatonomus miyakei Ochi & Araya 1992
Parascatonomus nilgiripolitus Ochi, Kon & Barclay 2017
Parascatonomus paotao Ochi, Kon & Barclay 2017
Parascatonomus paramairuu Ochi & Kon 2016
Parascatonomus politus (Paulian 1932)
Parascatonomus seminiasensis Ochi & Kon 2016
Parascatonomus shanicus Kabakov 2006
Parascatonomus sikkimpolitus Ochi, Kon & Barclay 2017
Parascatonomus vietnampolitus Ochi, Kon & Barclay 2017
Acanthonitis
Aliuscanthoniola
Allonitis
Anoctus
Anomiopsoides
Anonychonitis
Anoplodrepanus
Apotolamprus
Attavicinus
Ausmontins
Bdelyrus
Besourenga
Byrrhidium
Cambefortantus
Cambefortius
Canthon
Canthonidia
Catharsiocopris
Catharsius
Chalcocopris
Cheironitis
Clypeodrepanus
Copridaspidus
Copris
Coprophanaeus
Delopleurus
Deltochilum
Deltorhinum
Dendropaemon
Digitonthophagus
Diorygopyx
Dorbignyolus
Drepanocerus
Drepanoplatynus
Drogo
Dwesasilvasedis
Epidrepanus
Escarabaeus
Eudinopus
Euoniticellus
Eusaproecius
Frankenbergerius
Gilletellus
Glyphoderus
Gymnopleurus
Helictopleurus
Heliocopris
Heteroclitopus
Heteronitis
Holocephalus
Homocopris
Homocorpis
Hypocanthidium
Isocopris
Jossonthophagus
Latodrepanus
Liatongus
Martinezidium
Megathopa
Melanocanthon
Mentophilus
Microcopris
Mimonthophagus
Mnematidium
Monoplistes
Morettius
Neosaproecius
Ochicanthon
Oniticellus
Onitis
Onthophagus
Oxysternon
Pachylomera
Paracanthon
Paracryptocanthon
Paronthobium
Pedaria
Phaedotrogus
Pinacopodius
Pinacotarsus
Platyonitis
Pleuronitis
Pseudocopris
Pseudonthobium
Pseudopedaria
Pseudosaproecius
Scaptocnemis
Scarabaeus
Scatonomus
Scatrichus
Sceliages
Scybalocanthon
Silvinha
Streblopus
Sulcophanaeus
Temnoplectron
Theotimius
Trichillidium
Trichillum
Tropidonitis
178 additional siblings truncated for brevity.
See the
resource file
for a full list.
EOL Dynamic Hierarchy 1.1
Life
Cellular Organisms
Eukaryota
Opisthokonta
Metazoa
Bilateria
Protostomia
Ecdysozoa
Panarthropoda
Arthropoda
Pancrustacea
Hexapoda
Insecta
Pterygota
Neoptera
Endopterygota
Coleoptera
Polyphaga
Scarabaeiformia
Scarabaeoidea
Scarabaeidae
Parascatonomus
(this page)
Parascatonomus (Granulidorsum) aper (Sharp 1875)
Parascatonomus (Granulidorsum) dejongi (Krikken 1986)
Parascatonomus (Granulidorsum) hayashimasaoi (Ochi & Kon 1999)
Parascatonomus (Granulidorsum) koichii Ochi & Araya 1992
Parascatonomus (Granulidorsum) leytensis Ochi & Araya 1992
Parascatonomus (Granulidorsum) negrosensis Ochi & Araya 1992
Parascatonomus (Granulidorsum) pseudorudis Ochi & Araya 1992
Parascatonomus (Granulidorsum) quezonensis Ochi & Araya 1992
Parascatonomus (Granulidorsum) rudis (Sharp 1875)
Parascatonomus (Granulidorsum) sakamakii (Ochi, Kon & Kawahara 2010)
Parascatonomus (Granulidorsum) shizumui Ochi & Araya 1992
Parascatonomus (Granulidorsum) takedai Ochi & Araya 1992
Parascatonomus (Granulidorsum) takeshii Ochi & Araya 1992
Parascatonomus (Granulidorsum) yvescambeforti Ochi & Araya 1992
Parascatonomus (Necramator) accedens (Lansberge 1883)
Parascatonomus (Necramator) acuticollis (Gillet 1927)
Parascatonomus (Necramator) aerumnosus (Balthasar 1963)
Parascatonomus (Necramator) alexeevi (Tarasov, Krikken & Huijbregts 2010)
Parascatonomus (Necramator) anguicorius (Boucomont 1914)
Parascatonomus (Necramator) anitidus (Ochi & Kon 2005)
Parascatonomus (Necramator) aokii (Nomura 1976)
Parascatonomus (Necramator) atratus Kabakov 1983
Parascatonomus (Necramator) aurifex (Harold 1877)
Parascatonomus (Necramator) brendelli Ochi, Kon & Barclay 2008
Parascatonomus (Necramator) bundutuhanensis Ochi, Kon & Barclay 2008
Parascatonomus (Necramator) chaiyaphumensis Masumoto & Ochi 2014
Parascatonomus (Necramator) chapaensis (Kabakov 1983)
Parascatonomus (Necramator) chiangraiensis (Masumoto, Ochi & Hanboonsong 2007)
Parascatonomus (Necramator) damaki (Masumoto, Hanboonsong & Ochi 2002)
Parascatonomus (Necramator) danumcupreus (Krikken & Huijbregts 2009)
Parascatonomus (Necramator) diversiformis (Boucomont 1914)
Parascatonomus (Necramator) dux (Sharp 1875)
Parascatonomus (Necramator) formosanus (Gillet 1924)
Parascatonomus (Necramator) funebris (Boucomont 1919)
Parascatonomus (Necramator) gunsalami (Ochi & Kon 2005)
Parascatonomus (Necramator) horni (Balthasar 1935)
Parascatonomus (Necramator) igneus (Vigors 1825)
Parascatonomus (Necramator) itiokai (Ochi, Kon & Kishimoto-Yamada 2010)
Parascatonomus (Necramator) itoi (Nomura 1976)
Parascatonomus (Necramator) javacupreus Krikken & Huijbregts 2009
Parascatonomus (Necramator) javanitidus (Ochi, Kon & Hartini 2010)
Parascatonomus (Necramator) katoi (Ochi & Araya 1996)
Parascatonomus (Necramator) kikutai (Ochi & Kon 2005)
Parascatonomus (Necramator) kiuchianus (Masumoto, Yang & Ochi 2004)
Parascatonomus (Necramator) klapperichi (Balthasar 1952)
Parascatonomus (Necramator) kuatunensis (Balthasar 1942)
Parascatonomus (Necramator) laevicollis (Lansberge 1883)
Parascatonomus (Necramator) lakyim (Masumoto 1990)
Parascatonomus (Necramator) liewi (Ochi & Kon 2005)
Parascatonomus (Necramator) maesaensis Masumoto & Ochi 2014
Parascatonomus (Necramator) mairuu (Masumoto 1989)
Parascatonomus (Necramator) masahiroi (Masumoto, Ochi & Hanboonsong 2008)
Parascatonomus (Necramator) monticupreus (Krikken & Huijbregts 2009)
Parascatonomus (Necramator) munetoshii Masumoto, Ochi & Sakchoowong 2012
Parascatonomus (Necramator) murasakianus (Nomura 1976)
Parascatonomus (Necramator) muticifrons (Endrödi 1973)
Parascatonomus (Necramator) naviculifer (Boucomont 1919)
Parascatonomus (Necramator) niasensis (Boucomont 1914)
Parascatonomus (Necramator) nitidus (Waterhouse 1875)
Parascatonomus (Necramator) nusatenggaricus (Krikken & Huijbreghts 2008)
Parascatonomus (Necramator) peninsulocupreus Krikken & Huijbregts 2009
Parascatonomus (Necramator) phraoensis (Masumoto 1990)
Parascatonomus (Necramator) pilularius (Lansberge 1883)
Parascatonomus (Necramator) piyawati Masumoto, Ochi & Hanboonsong 2002
Parascatonomus (Necramator) poringensis (Ochi & Kon 2005)
Parascatonomus (Necramator) quaestus (Sharp 1875)
Parascatonomus (Necramator) rhombocephalus Kabakov 1983
Parascatonomus (Necramator) salakensis (Ochi, Kon & Hartini 2010)
Parascatonomus (Necramator) sarawacus (Harold 1877)
Parascatonomus (Necramator) satoi (Ochi, Shimada & Kon 2003)
Parascatonomus (Necramator) sayapensis (Ochi & Kon 2005)
Parascatonomus (Necramator) semiaureus (Lansberge 1883)
Parascatonomus (Necramator) semicupreus (Harold 1877)
Parascatonomus (Necramator) semifex (Krikken & Huijbregts 2008)
Parascatonomus (Necramator) serapiensis (Ochi, Kon & Barclay 2008)
Parascatonomus (Necramator) shibatai (Nakane 1960)
Parascatonomus (Necramator) striaticollis Kabakov 1983
Parascatonomus (Necramator) sumawacus (Krikken & Huijbregts 2008)
Parascatonomus (Necramator) taichii (Ochi, Kon & Barclay 2008)
Parascatonomus (Necramator) takahashii Masumoto, Ochi & Hanboonsong 2013
Parascatonomus (Necramator) takakui (Ochi, Kon & Hartini 2009)
Parascatonomus (Necramator) tamijii (Kon, Sakai & Ochi 2000)
Parascatonomus (Necramator) taominorui Masumoto & Ochi 2014
Parascatonomus (Necramator) thaitai (Masumoto, Ochi & Hanboonsong 2008)
Parascatonomus (Necramator) tricornis (Wiedemann 1823)
Parascatonomus (Necramator) trungboensis Kabakov 1983
Parascatonomus (Necramator) utsunomiyae (Masumoto, Ochi & Hanboonsong 2007)
Parascatonomus (Necramator) vasiljevi Kabakov 1983
Parascatonomus (Necramator) vitalisi (Boucomont 1921)
Parascatonomus (Necramator) vividus (Arrow 1907)
Parascatonomus (Peculiaricerus) anceyi (Boucomont 1921)
Parascatonomus (Peculiaricerus) egregius (Arrow 1907)
Parascatonomus (Pseudonthophagus) penicillatus (Harold 1879)
Parascatonomus (Pseudonthophagus) yukae (Masumoto, Ochi & Hanboonsong 2002)
Parascatonomus apilularius (Masumoto 1995)
Parascatonomus balikpapanensis Ochi & Kon 2016
Parascatonomus discedens (Sharp 1875)
Parascatonomus fujiokai (Ochi & Araya 1996)
Parascatonomus gosoli (Masumoto 1989)
Parascatonomus impar (Kabakov 2006)
Parascatonomus javapolitus Ochi, Kon & Barclay 2017
Parascatonomus laotianus (Boucomont 1919)
Parascatonomus maculatus (Fabricius 1801)
Parascatonomus miyakei Ochi & Araya 1992
Parascatonomus nilgiripolitus Ochi, Kon & Barclay 2017
Parascatonomus paotao Ochi, Kon & Barclay 2017
Parascatonomus paramairuu Ochi & Kon 2016
Parascatonomus politus (Paulian 1932)
Parascatonomus seminiasensis Ochi & Kon 2016
Parascatonomus shanicus Kabakov 2006
Parascatonomus sikkimpolitus Ochi, Kon & Barclay 2017
Parascatonomus vietnampolitus Ochi, Kon & Barclay 2017
Acanthonitis
Aliuscanthoniola
Allonitis
Anoctus
Anomiopsoides
Anonychonitis
Anoplodrepanus
Apotolamprus
Ateuchites
Attavicinus
Ausmontins
Bdelyrus
Besourenga
Byrrhidium
Cambefortantus
Cambefortius
Canthon
Canthonidia
Catharsiocopris
Catharsius
Chalcocopris
Cheironitis
Clypeodrepanus
Copridaspidus
Copris
Coprophanaeus
Delopleurus
Deltochilum
Deltorhinum
Dendropaemon
Digitonthophagus
Diorygopyx
Dorbignyolus
Drepanocerus
Drepanoplatynus
Drogo
Dwesasilvasedis
Epidrepanus
Escarabaeus
Eudinopus
Euoniticellus
Eusaproecius
Frankenbergerius
Glyphoderus
Gymnopleurus
Helictopleurus
Heliocopris
Heteroclitopus
Heteronitis
Holcorobeus
Holocephalus
Homocopris
Homocorpis
Hypocanthidium
Isocopris
Jossonthophagus
Latodrepanus
Liatongus
Martinezidium
Megathopa
Melanocanthon
Microcopris
Mimonthophagus
Mnematidium
Monoplistes
Morettius
Neosaproecius
Ochicanthon
Oniticellus
Onitis
Onthophagus
Oxysternon
Pachylomera
Paracanthon
Paracryptocanthon
Paronthobium
Pedaria
Phaedotrogus
Pinacopodius
Pinacotarsus
Platyonitis
Pleuronitis
Pseudocopris
Pseudonthobium
Pseudopedaria
Pseudosaproecius
Scaptocnemis
Scarabaeus
Scatonomus
Scatrichus
Sceliages
Scybalocanthon
Silvinha
Streblopus
Sulcophanaeus
Temnoplectron
Theotimius
Trichillidium
Trichillum
Tropidonitis
189 additional siblings truncated for brevity.
See the
resource file
for a full list.
EOL Dynamic Hierarchy Version 2.2.4
Parascatonomus
(this page)
Parascatonomus (Granulidorsum) aper (Sharp 1875)
Parascatonomus (Granulidorsum) dejongi (Krikken 1986)
Parascatonomus (Granulidorsum) hayashimasaoi (Ochi & Kon 1999)
Parascatonomus (Granulidorsum) koichii Ochi & Araya 1992
Parascatonomus (Granulidorsum) leytensis Ochi & Araya 1992
Parascatonomus (Granulidorsum) negrosensis Ochi & Araya 1992
Parascatonomus (Granulidorsum) pseudorudis Ochi & Araya 1992
Parascatonomus (Granulidorsum) quezonensis Ochi & Araya 1992
Parascatonomus (Granulidorsum) rudis (Sharp 1875)
Parascatonomus (Granulidorsum) sakamakii (Ochi, Kon & Kawahara 2010)
Parascatonomus (Granulidorsum) shizumui Ochi & Araya 1992
Parascatonomus (Granulidorsum) takedai Ochi & Araya 1992
Parascatonomus (Granulidorsum) takeshii Ochi & Araya 1992
Parascatonomus (Granulidorsum) yvescambeforti Ochi & Araya 1992
Parascatonomus (Necramator) accedens (Lansberge 1883)
Parascatonomus (Necramator) acuticollis (Gillet 1927)
Parascatonomus (Necramator) aerumnosus (Balthasar 1963)
Parascatonomus (Necramator) alexeevi (Tarasov, Krikken & Huijbregts 2010)
Parascatonomus (Necramator) anguicorius (Boucomont 1914)
Parascatonomus (Necramator) anitidus (Ochi & Kon 2005)
Parascatonomus (Necramator) aokii (Nomura 1976)
Parascatonomus (Necramator) atratus (Kabakov 1983)
Parascatonomus (Necramator) aurifex (Harold 1877)
Parascatonomus (Necramator) brendelli Ochi, Kon & Barclay 2008
Parascatonomus (Necramator) bundutuhanensis Ochi, Kon & Barclay 2008
Parascatonomus (Necramator) chaiyaphumensis Masumoto & Ochi 2014
Parascatonomus (Necramator) chapaensis (Kabakov 1983)
Parascatonomus (Necramator) chiangraiensis (Masumoto, Ochi & Hanboonsong 2007)
Parascatonomus (Necramator) damaki (Masumoto, Hanboonsong & Ochi 2002)
Parascatonomus (Necramator) danumcupreus (Krikken & Huijbregts 2009)
Parascatonomus (Necramator) diversiformis (Boucomont 1914)
Parascatonomus (Necramator) dux (Sharp 1875)
Parascatonomus (Necramator) formosanus (Gillet 1924)
Parascatonomus (Necramator) funebris (Boucomont 1919)
Parascatonomus (Necramator) gunsalami (Ochi & Kon 2005)
Parascatonomus (Necramator) horni (Balthasar 1935)
Parascatonomus (Necramator) igneus (Vigors 1825)
Parascatonomus (Necramator) itiokai (Ochi, Kon & Kishimoto-Yamada 2010)
Parascatonomus (Necramator) itoi (Nomura 1976)
Parascatonomus (Necramator) javacupreus Krikken & Huijbregts 2009
Parascatonomus (Necramator) javanitidus (Ochi, Kon & Hartini 2010)
Parascatonomus (Necramator) katoi (Ochi & Araya 1996)
Parascatonomus (Necramator) kikutai (Ochi & Kon 2005)
Parascatonomus (Necramator) kiuchianus (Masumoto, Yang & Ochi 2004)
Parascatonomus (Necramator) klapperichi (Balthasar 1952)
Parascatonomus (Necramator) kuatunensis (Balthasar 1942)
Parascatonomus (Necramator) laevicollis (Lansberge 1883)
Parascatonomus (Necramator) lakyim (Masumoto 1990)
Parascatonomus (Necramator) liewi (Ochi & Kon 2005)
Parascatonomus (Necramator) maesaensis Masumoto & Ochi 2014
Parascatonomus (Necramator) mairuu (Masumoto 1989)
Parascatonomus (Necramator) masahiroi (Masumoto, Ochi & Hanboonsong 2008)
Parascatonomus (Necramator) monticupreus (Krikken & Huijbregts 2009)
Parascatonomus (Necramator) munetoshii Masumoto, Ochi & Sakchoowong 2012
Parascatonomus (Necramator) murasakianus (Nomura 1976)
Parascatonomus (Necramator) muticifrons (Endrödi 1973)
Parascatonomus (Necramator) naviculifer (Boucomont 1919)
Parascatonomus (Necramator) niasensis (Boucomont 1914)
Parascatonomus (Necramator) nitidus (Waterhouse 1875)
Parascatonomus (Necramator) nusatenggaricus (Krikken & Huijbreghts 2008)
Parascatonomus (Necramator) peninsulocupreus Krikken & Huijbregts 2009
Parascatonomus (Necramator) phraoensis (Masumoto 1990)
Parascatonomus (Necramator) pilularius (Lansberge 1883)
Parascatonomus (Necramator) piyawati Masumoto, Ochi & Hanboonsong 2002
Parascatonomus (Necramator) poringensis (Ochi & Kon 2005)
Parascatonomus (Necramator) quaestus (Sharp 1875)
Parascatonomus (Necramator) rhombocephalus (Kabakov 1983)
Parascatonomus (Necramator) salakensis (Ochi, Kon & Hartini 2010)
Parascatonomus (Necramator) sarawacus (Harold 1877)
Parascatonomus (Necramator) satoi (Ochi, Shimada & Kon 2003)
Parascatonomus (Necramator) sayapensis (Ochi & Kon 2005)
Parascatonomus (Necramator) semiaureus (Lansberge 1883)
Parascatonomus (Necramator) semicupreus (Harold 1877)
Parascatonomus (Necramator) semifex (Krikken & Huijbregts 2008)
Parascatonomus (Necramator) serapiensis (Ochi, Kon & Barclay 2008)
Parascatonomus (Necramator) shibatai (Nakane 1960)
Parascatonomus (Necramator) striaticollis (Kabakov 1983)
Parascatonomus (Necramator) sumawacus (Krikken & Huijbregts 2008)
Parascatonomus (Necramator) taichii (Ochi, Kon & Barclay 2008)
Parascatonomus (Necramator) takahashii Masumoto, Ochi & Hanboonsong 2013
Parascatonomus (Necramator) takakui (Ochi, Kon & Hartini 2009)
Parascatonomus (Necramator) tamijii (Kon, Sakai & Ochi 2000)
Parascatonomus (Necramator) taominorui Masumoto & Ochi 2014
Parascatonomus (Necramator) thaitai (Masumoto, Ochi & Hanboonsong 2008)
Parascatonomus (Necramator) tricornis (Wiedemann 1823)
Parascatonomus (Necramator) trungboensis (Kabakov 1983)
Parascatonomus (Necramator) utsunomiyae (Masumoto, Ochi & Hanboonsong 2007)
Parascatonomus (Necramator) vasiljevi Kabakov 1992
Parascatonomus (Necramator) vitalisi (Boucomont 1921)
Parascatonomus (Necramator) vividus (Arrow 1907)
Parascatonomus (Peculiaricerus) anceyi (Boucomont 1921)
Parascatonomus (Peculiaricerus) egregius (Arrow 1907)
Parascatonomus (Pseudonthophagus) penicillatus (Harold 1879)
Parascatonomus (Pseudonthophagus) yukae (Masumoto, Ochi & Hanboonsong 2002)
Parascatonomus abramovi (Zinchenko 2018)
Parascatonomus apilularius (Masumoto 1995)
Parascatonomus balikpapanensis Ochi & Kon 2016
Parascatonomus discedens (Sharp 1875)
Parascatonomus fujiokai (Ochi & Araya 1996)
Parascatonomus gosoli (Masumoto 1989)
Parascatonomus impar (Kabakov 2006)
Parascatonomus javapolitus Ochi, Kon & Barclay 2017
Parascatonomus laotianus (Boucomont 1919)
Parascatonomus maculatus (Fabricius 1801)
Parascatonomus miyakei Ochi & Araya 1992
Parascatonomus nilgiripolitus Ochi, Kon & Barclay 2017
Parascatonomus paotao Ochi, Kon & Barclay 2017
Parascatonomus paramairuu Ochi & Kon 2016
Parascatonomus politus (Paulian 1932)
Parascatonomus seminiasensis Ochi & Kon 2016
Parascatonomus shanicus Kabakov 2006
Parascatonomus sikkimpolitus Ochi, Kon & Barclay 2017
Parascatonomus vietnampolitus Ochi, Kon & Barclay 2017
Acanthonitis
Aliuscanthoniola
Allonitis
Anoctus
Anomiopsoides
Anonychonitis
Anoplodrepanus
Apotolamprus
Attavicinus
Ausmontins
Bdelyrus
Besourenga
Byrrhidium
Cambefortantus
Cambefortius
Canthon
Canthonidia
Catharsiocopris
Catharsius
Chalcocopris
Cheironitis
Clypeodrepanus
Copridaspidus
Copris
Coprophanaeus
Delopleurus
Deltochilum
Deltorhinum
Dendropaemon
Digitonthophagus
Diorygopyx
Dorbignyolus
Drepanocerus
Drepanoplatynus
Drogo
Dwesasilvasedis
Epidrepanus
Escarabaeus
Eudinopus
Euoniticellus
Eusaproecius
Frankenbergerius
Gilletellus
Glyphoderus
Gymnopleurus
Helictopleurus
Heliocopris
Heteroclitopus
Heteronitis
Holocephalus
Homocopris
Homocorpis
Hypocanthidium
Isocopris
Jossonthophagus
Latodrepanus
Liatongus
Martinezidium
Megathopa
Melanocanthon
Mentophilus
Microcopris
Mimonthophagus
Mnematidium
Monoplistes
Morettius
Neosaproecius
Ochicanthon
Oniticellus
Onitis
Onthophagus
Oxysternon
Pachylomera
Paracanthon
Paracryptocanthon
Paronthobium
Pedaria
Phaedotrogus
Pinacopodius
Pinacotarsus
Platyonitis
Pleuronitis
Pseudocopris
Pseudonthobium
Pseudopedaria
Pseudosaproecius
Scaptocnemis
Scarabaeus
Scatonomus
Scatrichus
Sceliages
Scybalocanthon
Silvinha
Streblopus
Sulcophanaeus
Temnoplectron
Theotimius
Trichillidium
Trichillum
Tropidonitis
178 additional siblings truncated for brevity.
See the
resource file
for a full list.
GBIF classification
Animalia
Arthropoda
Insecta
Coleoptera
Scarabaeidae
Parascatonomus Paulian 1932
(this page)
Onthophagus miyakei Ochi 1992
Parascatonomus abramovi (Zinchenko 2018)
Parascatonomus accedens (Lansberge 1883)
Parascatonomus acuticollis (Gillet 1927)
Parascatonomus aerumnosus (Balthasar 1963)
Parascatonomus alexeevi (Tarasov, Krikken & Huijbregts 2010)
Parascatonomus anceyi (Boucomont 1921)
Parascatonomus anguicorius (Boucomont 1914)
Parascatonomus anitidus (Ochi & Kon 2005)
Parascatonomus aokii (Nomura 1976)
Parascatonomus aper (Sharp 1875)
Parascatonomus apilularius (Masumoto 1995)
Parascatonomus atratus (Kabakov 1983)
Parascatonomus aurifex (Harold 1877)
Parascatonomus balikpapanensis Ochi & Kon 2016
Parascatonomus brendelli Ochi, Kon & Barclay 2008
Parascatonomus bundutuhanensis Ochi, Kon & Barclay 2008
Parascatonomus chaiyaphumensis Masumoto & Ochi 2014
Parascatonomus chapaensis (Kabakov 1983)
Parascatonomus chiangraiensis (Masumoto, Ochi & Hanboonsong 2007)
Parascatonomus damaki (Masumoto, Hanboonsong & Ochi 2002)
Parascatonomus danumcupreus (Krikken & Huijbregts 2009)
Parascatonomus dejongi (Krikken 1986)
Parascatonomus discedens (Sharp 1875)
Parascatonomus diversiformis (Boucomont 1914)
Parascatonomus dux (Sharp 1875)
Parascatonomus egregius (Arrow 1907)
Parascatonomus formosanus (Gillet 1924)
Parascatonomus funebris (Boucomont 1919)
Parascatonomus gosoli (Masumoto 1989)
Parascatonomus gunsalami (Ochi & Kon 2005)
Parascatonomus hayashimasaoi (Ochi & Kon 1999)
Parascatonomus horni (Balthasar 1935)
Parascatonomus igneus (Vigors 1825)
Parascatonomus impar (Kabakov 2006)
Parascatonomus itiokai (Ochi, Kon & Kishimoto-Yamada 2010)
Parascatonomus itoi (Nomura 1976)
Parascatonomus javacupreus Krikken & Huijbregts 2009
Parascatonomus javanitidus (Ochi, Kon & Hartini 2010)
Parascatonomus javapolitus Ochi, Kon & Barclay 2017
Parascatonomus katoi (Ochi & Araya 1996)
Parascatonomus kiuchianus (Masumoto, Yang & Ochi 2004)
Parascatonomus klapperichi (Balthasar 1952)
Parascatonomus koichii Ochi & Araya 1992
Parascatonomus kuatunensis (Balthasar 1942)
Parascatonomus laevicollis (Lansberge 1883)
Parascatonomus lakyim (Masumoto 1990)
Parascatonomus laotianus (Boucomont 1919)
Parascatonomus leytensis Ochi & Araya 1992
Parascatonomus liewi (Ochi & Kon 2005)
Parascatonomus maculatus (Fabricius 1801)
Parascatonomus maesaensis Masumoto & Ochi 2014
Parascatonomus mairuu (Masumoto 1989)
Parascatonomus masahiroi (Masumoto, Ochi & Hanboonsong 2008)
Parascatonomus miyakei Ochi & Araya 1992
Parascatonomus monticupreus (Krikken & Huijbregts 2009)
Parascatonomus munetoshii Masumoto, Ochi & Sakchoowong 2012
Parascatonomus murasakianus (Nomura 1976)
Parascatonomus muticifrons (Endrödi 1973)
Parascatonomus naviculifer (Boucomont 1919)
Parascatonomus negrosensis Ochi & Araya 1992
Parascatonomus niasensis (Boucomont 1914)
Parascatonomus nilgiripolitus Ochi, Kon & Barclay 2017
Parascatonomus nitidus (Waterhouse 1875)
Parascatonomus nusatenggaricus (Krikken & Huijbreghts 2008)
Parascatonomus paotao Ochi, Kon & Barclay 2017
Parascatonomus paramairuu Ochi & Kon 2016
Parascatonomus penicillatus (Harold 1879)
Parascatonomus peninsulocupreus Krikken & Huijbregts 2009
Parascatonomus phraoensis (Masumoto 1990)
Parascatonomus pilularius (Lansberge 1883)
Parascatonomus piyawati Masumoto, Ochi & Hanboonsong 2002
Parascatonomus politus (Paulian 1932)
Parascatonomus poringensis (Ochi & Kon 2005)
Parascatonomus pseudorudis Ochi & Araya 1992
Parascatonomus quaestus (Sharp 1875)
Parascatonomus quezonensis Ochi & Araya 1992
Parascatonomus rhombocephalus (Kabakov 1983)
Parascatonomus sakamakii (Ochi, Kon & Kawahara 2010)
Parascatonomus salakensis (Ochi, Kon & Hartini 2010)
Parascatonomus sarawacus (Harold 1877)
Parascatonomus sayapensis (Ochi & Kon 2005)
Parascatonomus semiaureus (Lansberge 1883)
Parascatonomus semicupreus (Harold 1877)
Parascatonomus semifex (Krikken & Huijbregts 2008)
Parascatonomus seminiasensis Ochi & Kon 2016
Parascatonomus serapiensis (Ochi, Kon & Barclay 2008)
Parascatonomus shanicus Kabakov 2006
Parascatonomus shibatai (Nakane 1960)
Parascatonomus shizumui Ochi & Araya 1992
Parascatonomus sikkimpolitus Ochi, Kon & Barclay 2017
Parascatonomus striaticollis (Kabakov 1983)
Parascatonomus sumawacus (Krikken & Huijbregts 2008)
Parascatonomus taichii (Ochi, Kon & Barclay 2008)
Parascatonomus takahashii Masumoto, Ochi & Hanboonsong 2013
Parascatonomus takedai Ochi & Araya 1992
Parascatonomus takeshii Ochi & Araya 1992
Parascatonomus tamijii (Kon, Sakai & Ochi 2000)
Parascatonomus taominorui Masumoto & Ochi 2014
Parascatonomus thaitai (Masumoto, Ochi & Hanboonsong 2008)
Parascatonomus tricornis (Wiedemann 1823)
Parascatonomus trungboensis (Kabakov 1983)
Parascatonomus utsunomiyae (Masumoto, Ochi & Hanboonsong 2007)
Parascatonomus vasiljevi Kabakov 1992
Parascatonomus vietnampolitus Ochi, Kon & Barclay 2017
Parascatonomus vitalisi (Boucomont 1921)
Parascatonomus vividus (Arrow 1907)
Parascatonomus yukae (Masumoto, Ochi & Hanboonsong 2002)
Parascatonomus yvescambeforti Ochi & Araya 1992
Acanthonitis Janssens 1937
Afrodrepanus Krikken 2009
Agamopus Bates 1887
Amietina Cambefort 1981
Anoctus Sharp 1875
Anonychonitis Janssens 1950
Anoplosiagum Blanchard 1850
Aptenocanthon Matthews 1974
Aptychonitis Janssens 1937
Ateuchus Weber 1801
Attavicinus Philips & Bell 2008
Ausmontins Deschodt & Davis 2018
Baloghonthobium Paulian 1986
Bohepilissus Paulian 1975
Bolbites Harold 1868
Caccobiomorphus Balthasar 1964
Caccobius Thomson 1859
Caeconthobium Paulian 1984
Cambefortius Branco 1989
Canthochilum Chapin 1934
Canthonosoma Mac Leay 1871
Catharsiocopris Balthasar 1967
Cephalodesmius Westwood 1842
Cetonischema Reitter 1899
Cheironitis Lansberge 1875
Chiron Macleay 1819
Clypeodrepanus Krikken 2009
Copridaspidus Boucomont 1920
Coprophanaeus D' Olsoufieff 1924
Coptorhina Hope 1833
Ctilocephala Burmeister 1855
Degallieridium
Deltepilissus Silvério Pereira 1949
Dendropaemon Perty 1830
Diathermus Kraatz 1897
Dicranocara Frolov & Scholtz 2003
Dicronorhina Hope 1837
Dorbignyolus Branco 1989
Ennearabdus Van Lansberge 1874
Eucetonia Schoch 1894
Euglypta Mohnike 1873
Eusaproecius Branco 1989
Feeridium Vaz de Mello 2008
Garreta Janssens 1940
Glaphyrocanthon Martínez 1948
Glyphoderus Westwood 1837
Haematonotus Kraatz 1880
Hammondantus Cambefort 1978
Heterorrhina
Holocanthon Martínez & Pereira 1956
Hybosorites Nikolajev 1996
Lachnosterna Hope 1837
Leotrichillum
Lithoscarabaeus Nikolajev 1992
Lophorrhinides
Macroderes Westwood 1842
Macrophylla Hope 1837
Mecinonota Kraatz 1892
Megalorhina Westwood 1847
Melanocanthon Halffter 1958
Mentophilus Laporte de Castelnau 1840
Metacatharsius Paulian 1939
Micrelaphinis Schoch 1896
Miltotrogus Reitter 1902
Mimeoma Casey 1915
Mimonthophagus Balthasar 1963
Mnematium Macleay 1821
Nebulasilvius Deschodt & Scholtz 2008
Neosaproecius Branco 1990
Neptunides Thomson 1879
Nesosisyphus Vinson 1946
Nesovinsonia Martínez & Pereira 1959
Nunoidium
Onychothecus Boucomont 1912
Oxycetonia Arrow 1910
Oxygrylius Casey 1915
Pachylomera Kirby 1832
Pachysoma Macleay 1821
Paroniticellus Balthasar 1963
Parvuhowdenius Deschodt & Scholtz 2008
Peckolus Scholtz & Howden 1987
Pereiraidium
Phycocus Broun 1883
Pinacopodius Branco 1992
Pleuronitis Van Lansberge 1875
Proteroscarabaeus Grabau 1923
Pseudeuryomia Kraatz 1894
Pseudoapterogyna Martínez De La Escalera 1914
Pycnopanelus Arrow 1931
Rhinacosmus Kraatz 1895
Scarabaeus
Scybalophagus
Silvinha
Sisyphus Latreille 1807
Stenovalgus Kolbe 1892
Sukelus Branco 1992
Thaumastopeus Kraatz 1885
Tiniocellus Péringuey 1902
Trichiomorphus Bourgoin 1919
Xenocanthon Martínez 1952
258 additional siblings truncated for brevity.
See the
resource file
for a full list.
NCBI
root
cellular organisms
Eukaryota
Opisthokonta
Metazoa
Eumetazoa
Bilateria
Protostomia
Ecdysozoa
Panarthropoda
Arthropoda
Mandibulata
Pancrustacea
Hexapoda
Insecta
Dicondylia
Pterygota
Neoptera
Holometabola
Coleoptera
Polyphaga
Scarabaeiformia
Scarabaeoidea
Scarabaeidae
Scarabaeinae
Scarabaeinae incertae sedis
Parascatonomus
(this page)
Parascatonomus dux
Parascatonomus penicillatus
Parascatonomus rudis
Parascatonomus sarawacus
Parascatonomus semiaureus
Parascatonomus semicupreus
Anomiopus
Anonthobium
Anonychonitis
Aptychonitis
Bdelyropsis
Bdelyrus
Bolbites
Bubas
Caccobius
Canthodimorpha
Catharsius
Cheironitis
Cleptocaccobius
Coprophanaeus
Coptorhina
Cyptochirus
Delopleurus
Demarziella
Dendropaemon
Diabroctis
Dichotomius
Dicranocara
Digitonthophagus
Drepanocerus
Euoniticellus
Euonthophagus
Frankenbergerius
Gromphas
Helictopleurus
Heliocopris
Heteronitis
Heterosyphus
Hyalonthophagus
Janssensantus
Liatongus
Macroderes
Metacatharsius
Microcopris
Milichus
Neateuchus
Ochicanthon
Oniticellus
Onitis
Ontherus
Onthophagus
Oruscatus
Oxysternon
Pachysoma
Parachorius
Paracopris
Paraphytus
Paronthobium
Pedaria
Phalops
Phanaeus
Proagoderus
Pseudoarachnodes
Sarophorus
Scaptodera
Scatimus
Serrophorus
Sulcophanaeus
Tiniocellus
Tragiscus
Trichillum
Uroxys
Xinidium
wikidata_hierarchy tar gz
Biota
Eukaryota
Unikonta
Opisthokonta
Holozoa
Filozoa
Apoikozoa
Animalia
Eumetazoa
Bilateria
Protostomia
Ecdysozoa
Arthropoda
Hexapoda
Insecta
Dicondylia
Pterygota
Metapterygota
Neoptera
Eumetabola
Endopterygota
Coleoptera
Polyphaga
Scarabaeiformia
Scarabaeoidea
Scarabaeidae
Scarabaeinae
Onthophagini
Onthophagus
Parascatonomus
(this page)
Onthophagus angularis
Onthophagus asper
Onthophagus aspericollis
Onthophagus asperipennis
Onthophagus aspernatus
Onthophagus asperrimus
Onthophagus asperulus
Onthophagus astigma
Onthophagus athiensis
Onthophagus atricolor
Onthophagus atridorsis
Onthophagus atroaereus
Onthophagus atrofasciatus
Onthophagus atronitidus
Onthophagus atrosericeus
Onthophagus aureofuscus
Onthophagus aureopilosus
Onthophagus auriculatus
Onthophagus aurifex
Onthophagus australis
Onthophagus axillaris
Onthophagus aztecus
Onthophagus comperei
Onthophagus compositus
Onthophagus costatus
Onthophagus costiger
Onthophagus creber
Onthophagus cribellum
Onthophagus cribratus
Onthophagus cribripennis
Onthophagus curvispina
Onthophagus cyaneiceps
Onthophagus cyanellus
Onthophagus cyaneoniger
Onthophagus cyanochlorus
Onthophagus cyclographus
Onthophagus cynomysi
Onthophagus cyobioides
Onthophagus dama
Onthophagus dandalu
Onthophagus dapcauensis
Onthophagus dapitanensis
Onthophagus decedens
Onthophagus digitifer
Onthophagus discrepans
Onthophagus endroedii
Onthophagus epilamprus
Onthophagus frontalis
Onthophagus gaillardi
Onthophagus gajo
Onthophagus galeatus
Onthophagus ganalensis
Onthophagus gandju
Onthophagus gangeticus
Onthophagus gangulu
Onthophagus gazellinus
Onthophagus geelongensis
Onthophagus gemellatus
Onthophagus gemmatus
Onthophagus gestroi
Onthophagus gibber
Onthophagus gibberosus
Onthophagus gibbidorsis
Onthophagus gibbus
Onthophagus gidju
Onthophagus giraffa
Onthophagus glabratus
Onthophagus glasunovi
Onthophagus gonopygus
Onthophagus gracilipes
Onthophagus humboldti
Onthophagus illyricus
Onthophagus indosinicus
Onthophagus indutus
Onthophagus inelegans
Onthophagus ineptus
Onthophagus inermiceps
Onthophagus inermicollis
Onthophagus inflaticollis
Onthophagus infucatus
Onthophagus inops
Onthophagus laborans
Onthophagus mocquerysi
Onthophagus modestus
Onthophagus noctis
Onthophagus pilauco
Onthophagus pluton
Onthophagus quadripustulatus
Onthophagus queenslandicus
Onthophagus quinquedens
Onthophagus quinquetuberculatus
Onthophagus ragazzii
Onthophagus rotundicollis
Onthophagus schultzei
Onthophagus sideki
Onthophagus simplex
Onthophagus simpliciceps
Onthophagus simulator
Onthophagus sinuosicollis
Onthophagus sinuosus
2369 additional siblings truncated for brevity.
See the
resource file
for a full list.
wikidata_hierarchy tar gz
Biota
Eukaryota
Unikonta
Opisthokonta
Holozoa
Filozoa
Apoikozoa
Animalia
Eumetazoa
Bilateria
Protostomia
Ecdysozoa
Arthropoda
Hexapoda
Insecta
Dicondylia
Pterygota
Metapterygota
Neoptera
Eumetabola
Endopterygota
Coleoptera
Polyphaga
Scarabaeiformia
Scarabaeoidea
Scarabaeidae
Parascatonomus
(this page)
Parascatonomus balikpapanensis
Parascatonomus chiangraiensis
Parascatonomus koichii
Parascatonomus munetoshii
Parascatonomus nusatenggaricus
Parascatonomus paramairuu
Parascatonomus pseudoriekoae
Parascatonomus seminiasensis
Parascatonomus takahashii
Parascatonomus taominorui
Adoretus
Aegialiinae
Apterepilissus
Ateuchus
Aulonocneminae
Bradypodidium
Bubas
Canthon
Chiron
Chironinae
Chrysina
Cleptocaccobius
Cretoscarabaeus
Cyphochilus
Deltorhinum
Diabroctis
Dynamopodinae
Endroedyolus
Eremazinae
Euonthophagus
Eupoecila
Glyphoderus
Hellaserica
Heteronitis
Homocorpis
Isocopris
Janssensantus
Labroma
Larhodius
Lepanus
Lophodonitis
Macroderes
Macropanelus
Malagoniella
Martinezidium
Matthewsius
Megathopa
Melanocanthon
Mentophilus
Micrelaphinis
Milichus
Monteithocanthon
Nebulasilvius
Nesocanthon
Notopedaria
Onychothecus
Oruscatus
Outenikwanus
Pachylomera
Paracanthon
Paracryptocanthon
Paraphacosomoides
Paroniticellus
Paronitis
Parvuhowdenius
Peckolus
Pedaridium
Penalus
Pereiraidium
Phaedotrogus
Phanaeus
Pinacotarsus
Pleuronitis
Pseudignambia
Pseudocanthon
Pseudochironitis
Pseudocopris
Pseudonthobium
Pseudopedaria
Pseudosaproecius
Pseuduroxys
Pteronyx
Pycnopanelus
Rutelinae
Saphobiamorpha
Saphobius
Saprosites
Sauvagesinella
Scaptocnemis
Scatonomus
Scatrichus
Sceliages
Scybalocanthon
Scybalophagus
Sinapisoma
Stiptocnemis
Stiptotarsus
Streblopus
Sulcophanaeus
Sylvicanthon
Temnoplectron
Tesserodon
Tesserodoniella
Theotimius
Thyregis
Trichillidium
Trichius
Xinidium
Xynedria
Zonocopris
303 additional siblings truncated for brevity.
See the
resource file
for a full list.