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names in breadcrumbs
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Scarabaeidae
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Scientific Names
Preferred Names
Scarabaeidae
Family recognized by
EOL Dynamic Hierarchy April 2022
Reference taxon from EOL Dynamic Hierarchy Trunk
Cephaloserica
Subgenus recognized by
wikidata_hierarchy tar gz
Euchiridae
Family recognized by
GBIF classification
Scarabaeidae
Family recognized by
wikipedia 中文维基百科
,
wikipedia 日本語
,
wikipedia русскую Википедию
,
wikipedia NL
,
wikipedia POL
,
wikipedia VI
,
wikipedia CZ
,
wikipedia FI
,
wikipedia NO
,
wikipedia RO
,
wikipedia UK
,
wikipedia CA
,
wikipedia SV
,
wikipedia TR
,
EOL Dynamic Hierarchy 1.1
,
EOL Dynamic Hierarchy 1.1
,
wikipedia ID
,
wikipedia EU
,
wikipedia AST
,
wikipedia AZ
,
wikipedia LT
,
wikipedia ET
,
wikipedia hr Croatian
,
wikipedia DA
,
wikipedia LA
,
wikipedia LA
,
wikipedia MS
,
wikipedia SK
,
wikipedia NN
,
wikipedia LV
,
wikipedia IS
,
wikipedia SL
,
wikipedia HSB
,
wikipedia GA
,
NCBI
,
wikipedia EO
,
Biopix Nature Photos
,
wikipedia emerging languages
,
wikipedia emerging_languages
,
wikipedia MIN
,
wikipedia gl Galician
,
GBIF classification
,
NCBI GGBN GBIF BHL BOLDS data coverage
,
Mozambique Species List
,
wikidata_hierarchy tar gz
,
wikipedia DE
,
wikipedia ES
,
wikipedia IT
,
wikipedia PT
,
wikipedia FR
,
wikipedia 한국어 위키백과
,
wikipedia EN
,
Wikidata Common Names
,
Wikimedia Commons
,
Barcode of Life Data Systems
,
Global Biotic Interactions
,
Global Biotic Interactions
,
Global Biotic Interactions
,
Global Biotic Interactions
,
iNaturalist data coverage
,
Wikipedia (inferred records)
,
GBIF data coverage
,
Dynamic Hierarchy Version 2.2.3
,
GGBN data coverage
,
wikipedia BR
, and
EOL Dynamic Hierarchy Version 2.2.4
Scarabaeidae
Recognized by
Eswatini Species List
,
CalPhotos
,
University of Alberta Museums
,
Natural History Museum Species of the day
,
INBio
,
IABIN
,
Ecomare
,
Moorea Biocode
,
Algeria Species List
,
Andorra Species List
,
Angola Species List
,
Armenia Species List
,
Austria Species List
,
Belarus Species List
,
Australia Species List
,
Belize Species List
,
Bhutan Species List
,
Belgium Species List
,
Bolivia Species List
,
Bosnia and Herzegovina Species List
,
Botswana Species List
,
Brazil Species List
,
Brunei Species List
,
Bulgaria Species List
,
Burkina Faso Species List
,
Burundi Species List
,
Cameroon Species List
,
Canada Species List
,
Chad Species List
,
Chile Species List
,
China Species List
,
Colombia Species List
,
Costa Rica Species List
,
Croatia Species List
,
Czech Republic Species List
,
Democratic Republic of the Congo Species List
,
Denmark Species List
,
Dominican Republic Species List
,
Ecuador Species List
,
El Salvador Species List
,
Estonia Species List
,
Ethiopia Species List
,
Finland Species List
,
France Species List
,
French Guiana Species List
,
Gabon Species List
,
Georgia Species List
,
Germany Species List
,
Ghana Species List
,
Greece Species List
,
Guam Species List
,
Guatemala Species List
,
Guernsey Species List
,
Guinea Species List
,
Guyana Species List
,
Haiti Species List
,
Hong Kong Species List
,
Hungary Species List
,
India Species List
,
Iran Species List
,
Iraq Species List
,
Israel Species List
,
Italy Species List
,
Ivory Coast Species List
,
Jamaica Species List
,
Kazakhstan Species List
,
Kenya Species List
,
Kosovo Species List
,
Latvia Species List
,
Lesotho Species List
,
Liberia Species List
,
Liechtenstein Species List
,
Lithuania Species List
,
Luxembourg Species List
,
Madagascar Species List
,
Malawi Species List
,
Malaysia Species List
,
Maldives Species List
,
Mariana Islands Species List
,
Mauritania Species List
,
Montenegro Species List
,
Morocco Species List
,
Namibia Species List
,
Nepal Species List
,
New Caledonia Species List
,
New Zealand Species List
,
Nicaragua Species List
,
Niger Species List
,
Nigeria Species List
,
Norway Species List
,
Pakistan Species List
,
Panama Species List
,
Papua New Guinea Species List
,
Peru Species List
,
Paraguay Species List
,
Philippines Species List
,
Poland Species List
,
Portugal Species List
,
Puerto Rico Species List
,
Republic of the Congo Species List
,
Romania Species List
,
Russia Species List
,
Rwanda Species List
,
Saudi Arabia Species List
,
Senegal Species List
,
Serbia Species List
,
Seychelles Species List
,
Singapore Species List
,
Slovakia Species List
,
Slovenia Species List
,
Somalia Species List
,
South Korea Species List
,
South Africa Species List
,
Spain Species List
,
Sudan Species List
,
Suriname Species List
,
Sweden Species List
,
Switzerland Species List
,
Syria Species List
,
Taiwan Species List
,
Tanzania Species List
,
Thailand Species List
,
The Bahamas Species List
,
The Netherlands Species List
,
Sri Lanka Species List
,
Turkey Species List
,
Uganda Species List
,
Ukraine Species List
,
United Arab Emirates Species List
,
United Kingdom Species List
,
Venezuela Species List
,
Vietnam Species List
,
Uruguay Species List
,
Uzbekistan Species List
,
Zambia Species List
,
Zimbabwe Species List
,
Argentina Species List
,
Japan Species List
,
English Vernaculars for Landmark Taxa
,
Indonesia Species List
,
Mexico Species List
,
Arabian Sea Species List
,
Gulf of Guinea Species List
,
Indian Ocean Species List
,
Mozambique Channel Species List
,
North Atlantic Species List
,
North Pacific Species List
,
South Atlantic Species List
,
South Pacific Species List
,
Furui 2020
,
Animal Diversity Web
,
OBIS environmental data records
,
GBIF national node type records Brazil
,
GBIF nat'l node type records Netherlands
,
GBIF national node type records Germany
,
United States Species List
, and
Flickr BHL
Scarabaeidae Latreille 1802
Family recognized by
Catalogue of Life 26 March 2024
and
Integrated Taxonomic Information System (ITIS) Taxonomic Hierarchy
Alternative Names
? baissensis Nikritin, 1977
Doubtful according to
GBIF classification
? labreae Pierce, 1946
Doubtful according to
GBIF classification
? leptoscelis Nikritin, 1977
Doubtful according to
GBIF classification
? oblongus Ponomarenko, 1990
Doubtful according to
GBIF classification
Acanthoprotaetia Miksic 1987
Doubtful according to
GBIF classification
Acanthosternum Philippi 1861
Doubtful according to
GBIF classification
Aceratocrates Ohaus 1910
Doubtful according to
GBIF classification
Achromisetes Kraatz 1880
Doubtful according to
GBIF classification
Actinophorus Creutzer 1799
Doubtful according to
GBIF classification
Acylochiloides Martinez 1975
Doubtful according to
GBIF classification
Adonides Thomson 1880
Doubtful according to
GBIF classification
Adoretus Laporte 1840
Doubtful according to
GBIF classification
Aegialatis Kono 1938
Doubtful according to
GBIF classification
Aemilius Le Moult 1939
Doubtful according to
GBIF classification
Aethiartrogus Decelle 1982
Doubtful according to
GBIF classification
Afrocanthon Müller 1941
Doubtful according to
GBIF classification
Afrocloeotus
Doubtful according to
GBIF classification
Afrodiasticus Pittino & Mariani 1986
Doubtful according to
GBIF classification
Afroharoldius Janssens 1949
Doubtful according to
GBIF classification
Aganhyboma Kolbe 1893
Doubtful according to
GBIF classification
Agnathocera Arrow 1922
Doubtful according to
GBIF classification
Alcidiosoma
Doubtful according to
GBIF classification
Aleiantus Olsoufieff 1947
Doubtful according to
GBIF classification
Aleurostictus Kirby 1827
Doubtful according to
GBIF classification
Alleurostictus
Doubtful according to
GBIF classification
Allophileurus
Doubtful according to
GBIF classification
Altonthophagus Kabakov 1990
Doubtful according to
GBIF classification
Amaecylius Huchet 2000
Doubtful according to
GBIF classification
Amalus Mulsant & Rey 1870
Doubtful according to
GBIF classification
Amartinezus Özdikmen 2009
Doubtful according to
GBIF classification
Amaurodes Westwood 1844
Doubtful according to
GBIF classification
Amauroleucocelis Bourgoin 1913
Doubtful according to
GBIF classification
Ammaecius
Doubtful according to
GBIF classification
Amphibolonarzon Gistl 1848
Doubtful according to
GBIF classification
Amphimallina Reitter 1905
Doubtful according to
GBIF classification
Amphimallon Berthold 1827
Doubtful according to
GBIF classification
Amphionthophagus Martin Piera & Zunino 1983
Doubtful according to
GBIF classification
Amphistous Lansberge 1871
Doubtful according to
GBIF classification
Amuarina
Doubtful according to
GBIF classification
Anacamphtorrhina Blanchard 1842
Doubtful according to
GBIF classification
Anachalcos Hope 1837
Doubtful according to
GBIF classification
Anaspilus Kolbe 1892
Doubtful according to
GBIF classification
Anatochilus Péringuey 1907
Doubtful according to
GBIF classification
Anatrinodia Casey 1915
Doubtful according to
GBIF classification
Ancistroma Curtis 1835
Doubtful according to
GBIF classification
Anectoma Gerstaecker 1883
Doubtful according to
GBIF classification
Anepsiovalgus Kolbe 1904
Doubtful according to
GBIF classification
Aneurystypus Blackburn 1888
Doubtful according to
GBIF classification
Aneustypus
Doubtful according to
GBIF classification
Anisoplia Fischer 1824
Doubtful according to
GBIF classification
Anisoplia Schönherr 1817
Doubtful according to
GBIF classification
Anomala Deshayes
Doubtful according to
GBIF classification
Anomalocera Westwood 1842
Doubtful according to
GBIF classification
Anomalochilus Blanchard 1850
Doubtful according to
GBIF classification
Anomocaulus Fairmaire 1881
Doubtful according to
GBIF classification
Anonetus Erichson 1847
Doubtful according to
GBIF classification
Anoplogomphus Prell 1912
Doubtful according to
GBIF classification
Anopsoistes
Doubtful according to
GBIF classification
Anthracopharis Schoch 1895
Doubtful according to
GBIF classification
Antillacanthon Vulcano & Pereira 1966
Doubtful according to
GBIF classification
Apanesthes Kraatz 1880
Doubtful according to
GBIF classification
Aphelorhina
Doubtful according to
GBIF classification
Aphelorrhina Westwood 1843
Doubtful according to
GBIF classification
Aphodiocopris Arrow 1920
Doubtful according to
GBIF classification
Aphodius
Doubtful according to
GBIF classification
Aphodus
Doubtful according to
GBIF classification
Aphonoproctus Kolbe 1905
Doubtful according to
GBIF classification
Aphonoprocus
Doubtful according to
GBIF classification
Aplasta Schaum 1845
Doubtful according to
GBIF classification
Aplodema Blanchard 1850
Doubtful according to
GBIF classification
Apohdius
Doubtful according to
GBIF classification
Apterocanthon
Doubtful according to
GBIF classification
Apterogyna Reiche 1882
Proparte synonym according to
GBIF classification
Apterogyna Reitter 1902
Doubtful according to
GBIF classification
Argyripia
Doubtful according to
GBIF classification
Argyrophegges Kraatz 1895
Doubtful according to
GBIF classification
Arrowianella Paulian 1933
Doubtful according to
GBIF classification
Arrowiellus Balthasar 1933
Doubtful according to
GBIF classification
Articephala Moser 1920
Doubtful according to
GBIF classification
Asserador Maunders 1852
Doubtful according to
GBIF classification
Astaborus Thomson 1859
Doubtful according to
GBIF classification
Astaenomochus
Doubtful according to
GBIF classification
Astaenosiagum Martínez 1957
Doubtful according to
GBIF classification
Astenorella
Doubtful according to
GBIF classification
Astenorhella Westwood 1873
Doubtful according to
GBIF classification
Astrocara
Doubtful according to
GBIF classification
Ataenia Schoch 1895
Doubtful according to
GBIF classification
Ataenioides Petrovitz 1973
Doubtful according to
GBIF classification
Atrichius Gillet 1907
Doubtful according to
GBIF classification
Atropinota Heller 1923
Doubtful according to
GBIF classification
Aulacium Reiche 1841
Doubtful according to
GBIF classification
Aulanota Moser 1924
Doubtful according to
GBIF classification
Aurelia Thomson 1880
Doubtful according to
GBIF classification
Australommoecius
Doubtful according to
GBIF classification
Autoprotaetia Mikšic 1965
Doubtful according to
GBIF classification
Baesia Burmeister 1842
Doubtful according to
GBIF classification
Baloghonthoboum Paulian 1986
Doubtful according to
GBIF classification
Barratus
Doubtful according to
GBIF classification
Barybas Burmeister 1855
Doubtful according to
GBIF classification
Bdelyopsis
Doubtful according to
GBIF classification
Belloanopsiostes Vaz de Mello 1996
Doubtful according to
GBIF classification
Bezdekia Özdikmen & Demir 2008
Doubtful according to
GBIF classification
Bielliana Baraud 1980
Doubtful according to
GBIF classification
Bifurcanomala Kim 1998
Doubtful according to
GBIF classification
Bislardiana Antoine 2003
Doubtful according to
GBIF classification
Bisoroclita Krikken 1982
Doubtful according to
GBIF classification
Blapephorus
Doubtful according to
GBIF classification
Boreocanthon Halffter 1958
Doubtful according to
GBIF classification
Bothrothorax Kraatz 1900
Doubtful according to
GBIF classification
Boucomontius d'Olsoufieff 1923
Doubtful according to
GBIF classification
Bourgoinigora Paulian 1994
Doubtful according to
GBIF classification
Brachymitra Kolbe 1904
Doubtful according to
GBIF classification
Brachymys Thomson 1858
Doubtful according to
GBIF classification
Brachytricha Reiche 1871
Doubtful according to
GBIF classification
Brenskiella Berg 1898
Doubtful according to
GBIF classification
Bryantella Britton 1957
Doubtful according to
GBIF classification
Byrromorpha Blackburn 1892
Doubtful according to
GBIF classification
Caccobius
Doubtful according to
GBIF classification
Caccocnemus
Doubtful according to
GBIF classification
Cacochroa Kraatz 1880
Doubtful according to
GBIF classification
Caelontherus Genier 1996
Doubtful according to
GBIF classification
Calocampsiura Miksic 1987
Doubtful according to
GBIF classification
Calochroea Schoch 1898
Doubtful according to
GBIF classification
Caloglycyphana Miksic 1968
Doubtful according to
GBIF classification
Calopotosia Reitter 1899
Doubtful according to
GBIF classification
Caloprotaetia Mikšic 1963
Doubtful according to
GBIF classification
Calysoryctes Howden 1970
Doubtful according to
GBIF classification
Camelonotus Fairmaire 1883
Doubtful according to
GBIF classification
Campulipes
Doubtful according to
GBIF classification
Campytrochalus Brenske 1900
Doubtful according to
GBIF classification
Canthomoechus Pereira & Martínez 1960
Doubtful according to
GBIF classification
Captomia Burmeister 1842
Doubtful according to
GBIF classification
Captomiopsis Pouillaude 1919
Doubtful according to
GBIF classification
Carolina Thomson 1880
Doubtful according to
GBIF classification
Cassolus Sharp 1875
Doubtful according to
GBIF classification
Celebesia Alexis & Delpont 1998
Doubtful according to
GBIF classification
Celebesiana Alexis & Delpont 2002
Doubtful according to
GBIF classification
Centrocheilus Krikken 1976
Doubtful according to
GBIF classification
Cephalocosmus Kraatz 1895
Doubtful according to
GBIF classification
Cephaloserica Brenske 1900
Doubtful according to
GBIF classification
Chacocopris
Doubtful according to
GBIF classification
Chaetodermatina Heller 1921
Doubtful according to
GBIF classification
Chaetopisthides Wasmann 1918
Doubtful according to
GBIF classification
Chaetoserica Brenske 1896
Doubtful according to
GBIF classification
Chalcoprotaetia Mikšic 1963
Doubtful according to
GBIF classification
Chapincanthon Vulcano & Pereira 1966
Doubtful according to
GBIF classification
Chasmatopterus Latreille 1825
Doubtful according to
GBIF classification
Chasmodia Macleay 1819
Doubtful according to
GBIF classification
Chejuanomala Kim 1998
Doubtful according to
GBIF classification
Chelorrhina Burmeister 1842
Doubtful according to
GBIF classification
Chilamblys Kraatz 1880
Doubtful according to
GBIF classification
Chirinda Péringuey 1907
Doubtful according to
GBIF classification
Chiriquiba
Doubtful according to
GBIF classification
Chironitis Bedel 1892
Doubtful according to
GBIF classification
Chrysoliocola Alexis & Delpont 1998
Doubtful according to
GBIF classification
Chtonobius
Doubtful according to
GBIF classification
Cineretis Schürhoff 1937
Doubtful according to
GBIF classification
Ciocotalpa Saylor 1940
Doubtful according to
GBIF classification
Cirrhospila Kraatz 1890
Doubtful according to
GBIF classification
Cjaradria
Doubtful according to
GBIF classification
Clineterocera
Doubtful according to
GBIF classification
Clitocephala Burmeister 1855
Doubtful according to
GBIF classification
Clydonophorus Kraatz 1899
Doubtful according to
GBIF classification
Clysterius Endrödi 1963
Doubtful according to
GBIF classification
Coelosis Kirby & Spence 1828
Doubtful according to
GBIF classification
Coelothorax Ancey 1880
Doubtful according to
GBIF classification
Colorhinus Erichson 1841
Doubtful according to
GBIF classification
Colpomorpha Szito 1994
Doubtful according to
GBIF classification
Coniotrogus Brenske 1903
Doubtful according to
GBIF classification
Copris Müller 1764
Doubtful according to
GBIF classification
Coprobius Latreille 1829
Doubtful according to
GBIF classification
Coprocanthon Martinez 1950
Doubtful according to
GBIF classification
Coproechus Reiche 1841
Doubtful according to
GBIF classification
Coprophanaeoides Edmonds 1972
Doubtful according to
GBIF classification
Coprophaneus
Doubtful according to
GBIF classification
Coprophnaeoides
Doubtful according to
GBIF classification
Coptorrhina
Doubtful according to
GBIF classification
Coquerelia Kraatz 1880
Doubtful according to
GBIF classification
Corminus Burmeister 1855
Doubtful according to
GBIF classification
Coronoserica Brenske 1900
Doubtful according to
GBIF classification
Corvicoana Strand 1934
Doubtful according to
GBIF classification
Coryphaeus Gistl 1847
Doubtful according to
GBIF classification
Coryphocera Burmeister 1842
Doubtful according to
GBIF classification
Cosmiomorphomima Miksic 1976
Doubtful according to
GBIF classification
Cosmiophaenia
Doubtful according to
GBIF classification
Cosmiorrhina Bourgoin 1918
Doubtful according to
GBIF classification
Cotinorrhina Schoch 1895
Doubtful according to
GBIF classification
Crecotoma Nikolajev 2002
Doubtful according to
GBIF classification
Cremastodus Solier 1851
Doubtful according to
GBIF classification
Crematochila
Doubtful according to
GBIF classification
Criniflava Goodrich 1966
Doubtful according to
GBIF classification
Cryphaeobius Kraatz 1882
Doubtful according to
GBIF classification
Cryptodontus Burlini 1969
Doubtful according to
GBIF classification
Cryptoryctes Carne 1957
Doubtful according to
GBIF classification
Crytocanthon Burmeister 1846
Doubtful according to
GBIF classification
Ctenotis Burmeister 1855
Doubtful according to
GBIF classification
Cyclocephala Latreille 1829
Doubtful according to
GBIF classification
Cyclophorellus Krikken 1982
Doubtful according to
GBIF classification
Cyclotrogus Wasmann 1918
Doubtful according to
GBIF classification
Cyphonoxia Reitter 1889
Doubtful according to
GBIF classification
Cyrrhochroea Kraatz 1900
Doubtful according to
GBIF classification
Daedycorrhina Bates 1880
Doubtful according to
GBIF classification
Daemonoplus Thomson 1858
Doubtful according to
GBIF classification
Damonplus
Doubtful according to
GBIF classification
Deltorrhinum Gemminger & Harold 1869
Doubtful according to
GBIF classification
Demodema Blanchard 1850
Doubtful according to
GBIF classification
Deronitis Arrow 1933
Doubtful according to
GBIF classification
Diasticus Mulsant 1842
Doubtful according to
GBIF classification
Dibolus Endrodi 1976
Doubtful according to
GBIF classification
Dicheloscema Moser 1924
Doubtful according to
GBIF classification
Dicranocephalus Hope 1831
Doubtful according to
GBIF classification
Dimyxus Fairmaire 1893
Doubtful according to
GBIF classification
Dineterogomphus Prell 1912
Doubtful according to
GBIF classification
Dineterohgomphus
Doubtful according to
GBIF classification
Dinonota Kraatz 1900
Doubtful according to
GBIF classification
Dinorhina Lacordaire 1856
Doubtful according to
GBIF classification
Diphydactylus Thomson 1858
Doubtful according to
GBIF classification
Diploeida Péringuey 1907
Doubtful according to
GBIF classification
Diplognathoides Holm & Perissinotto 2004
Doubtful according to
GBIF classification
Doedycorhina Bates 1888
Doubtful according to
GBIF classification
Dorcus Gistl 1847
Doubtful according to
GBIF classification
Doryscelis Burmeister 1842
Doubtful according to
GBIF classification
Drepanellus Barbero, Palestrini & Roggero 2009
Doubtful according to
GBIF classification
Dynastes Kirby 1825
Doubtful according to
GBIF classification
Dysphanocheila Blackburn 1898
Doubtful according to
GBIF classification
Dyspilophora Kraatz 1880
Doubtful according to
GBIF classification
Elaphoides Schoch 1894
Doubtful according to
GBIF classification
Elassochiton Kolbe 1895
Doubtful according to
GBIF classification
Endreodyolus
Doubtful according to
GBIF classification
Endrodius
Doubtful according to
GBIF classification
Endroedia Mikšic 1958
Doubtful according to
GBIF classification
Energetia Dalle Torre 1912
Doubtful according to
GBIF classification
Enicotarsus Laporte 1831
Doubtful according to
GBIF classification
Entyopsis Kolbe 1894
Doubtful according to
GBIF classification
Eoalleucosma Antoine 1989
Doubtful according to
GBIF classification
Eoheterorrhina Miksic 1974
Doubtful according to
GBIF classification
Eotrochalus Burgeon 1943
Doubtful according to
GBIF classification
Ephirinus
Doubtful according to
GBIF classification
Epichalcoplethis Bates 1904
Doubtful according to
GBIF classification
Epicometes
Doubtful according to
GBIF classification
Epicometis Burmeister 1842
Doubtful according to
GBIF classification
Epilissus Blanchard 1845
Doubtful according to
GBIF classification
Epionitis Balthasar 1942
Doubtful according to
GBIF classification
Eremonthophagus Zunino 1979
Doubtful according to
GBIF classification
Eremoryctes
Doubtful according to
GBIF classification
Estenomenus Faldermann 1835
Doubtful according to
GBIF classification
Esymaphodius
Doubtful according to
GBIF classification
Eubarybas Gutiérrez 1952
Doubtful according to
GBIF classification
Eucampsiura Miksic 1987
Doubtful according to
GBIF classification
Eucanthidium Martinez & Halffter 1986
Doubtful according to
GBIF classification
Eucatheta Fairmaire 1894
Doubtful according to
GBIF classification
Euchroa Gistl 1847
Doubtful according to
GBIF classification
Euchromoplia Medvedev 1952
Doubtful according to
GBIF classification
Eucoelosis Iannuzzi & Marinoni 1995
Doubtful according to
GBIF classification
Eucymophorus Schein 1954
Doubtful according to
GBIF classification
Euglycyphana Miksic 1968
Doubtful according to
GBIF classification
Euhyboma Kolbe 1893
Doubtful according to
GBIF classification
Euligyrus Casey 1915
Doubtful according to
GBIF classification
Eumimela Kraatz 1881
Doubtful according to
GBIF classification
Eupotosia Miksic 1954
Doubtful according to
GBIF classification
Euprigenia Miksic 1974
Doubtful according to
GBIF classification
Euprochyta Fairmaire 1899
Doubtful according to
GBIF classification
Euprotaetia Mikšic 1963
Doubtful according to
GBIF classification
Eurycorythoderus Wasmann 1918
Doubtful according to
GBIF classification
Euryderus Gemminger & Harold 1869
Doubtful according to
GBIF classification
Eurynomia
Doubtful according to
GBIF classification
Euryphylla Lacordaire 1856
Doubtful according to
GBIF classification
Eurypodea Klages 1906
Doubtful according to
GBIF classification
Eurysternodes Martinez 1988
Doubtful according to
GBIF classification
Eutheola
Doubtful according to
GBIF classification
Eutlesthes
Doubtful according to
GBIF classification
Excisivalgus Endrödi 1952
Doubtful according to
GBIF classification
Fairmairiana Alexis & Delpont 1998
Doubtful according to
GBIF classification
Ferreiria
Doubtful according to
GBIF classification
Finkia Mikšic 1965
Doubtful according to
GBIF classification
Formicdubius Philips & Scholtz 2000
Doubtful according to
GBIF classification
Fortishybosorus Yan, Bai & Ren 2013
Doubtful according to
GBIF classification
Foveopotosia Mikšic 1957
Doubtful according to
GBIF classification
Francmonrosia Pereira & Martinez 1960
Doubtful according to
GBIF classification
Freyus Balthasar 1958
Doubtful according to
GBIF classification
Furconthophagus Zunino 1979
Doubtful according to
GBIF classification
Gama Blanchard 1850
Doubtful according to
GBIF classification
Gastrohoplus Moser 1921
Doubtful according to
GBIF classification
Geminigora Paulian 1994
Doubtful according to
GBIF classification
Genuchinus Kirby 1825
Doubtful according to
GBIF classification
Genyoschiza Moser 1917
Doubtful according to
GBIF classification
Geocanthon Pereira & Martínez 1956
Doubtful according to
GBIF classification
Ghania Endrodi 1973
Doubtful according to
GBIF classification
Glaphyocanthon
Doubtful according to
GBIF classification
Glaraphodius Balthasar 1941
Doubtful according to
GBIF classification
Glaucocelis Kraatz 1897
Doubtful according to
GBIF classification
Glycyphaniola Miksic 1968
Doubtful according to
GBIF classification
Glyphiderus Westwood 1838
Doubtful according to
GBIF classification
Glyphoderus Agassiz 1846
Doubtful according to
GBIF classification
Glyptoterus
Doubtful according to
GBIF classification
Goetzia Mikšic 1963
Doubtful according to
GBIF classification
Goliata
Doubtful according to
GBIF classification
Goliathinus Westwood 1874
Doubtful according to
GBIF classification
Gomphas Brullé 1834
Doubtful according to
GBIF classification
Goniocanthon Pereira & Martínez 1956
Doubtful according to
GBIF classification
Gonoprotaetia Mikšic 1965
Doubtful according to
GBIF classification
Grebennikovius
Doubtful according to
GBIF classification
Grebennikovius Mlambo, Scholtz & Deschodt 2019
Doubtful according to
GBIF classification
Grypnocnemis
Doubtful according to
GBIF classification
Gymnetina Kirby 1827
Doubtful according to
GBIF classification
Gyronotus Lansberge 1874
Doubtful according to
GBIF classification
Hadrocerus Guérin-Ménéville 1838
Doubtful according to
GBIF classification
Haplonycha Agassiz 1846
Doubtful according to
GBIF classification
Haroldostes Paulian 1982
Doubtful according to
GBIF classification
Helionica Thomson 1881
Doubtful according to
GBIF classification
Hemicephalocosmus Miksic 1974
Doubtful according to
GBIF classification
Hemicoryphocera Miksic 1974
Doubtful according to
GBIF classification
Hemiopta Bedel 1896
Doubtful according to
GBIF classification
Hemiphaedimus Miksic 1972
Doubtful according to
GBIF classification
Hemipharia
Doubtful according to
GBIF classification
Hemiprigenia Miksic 1974
Doubtful according to
GBIF classification
Hemiserica Brenske 1894
Doubtful according to
GBIF classification
Hemitheodosia Sakai 1994
Doubtful according to
GBIF classification
Hemycirtus Reiche 1860
Doubtful according to
GBIF classification
Hemycorythoderus
Doubtful according to
GBIF classification
Heptophyllla Motschulsky 1857
Doubtful according to
GBIF classification
Hercites Burmeister 1855
Doubtful according to
GBIF classification
Heterelaphinis Antoine 2002
Doubtful according to
GBIF classification
Heteroateuchus Paulian 1935
Doubtful according to
GBIF classification
Heterocotinus Martinez 1948
Doubtful according to
GBIF classification
Heteroglycyphana Miksic 1968
Doubtful according to
GBIF classification
Heteronychidius Paulian 1954
Doubtful according to
GBIF classification
Heteropodia Schürhoff 1933
Doubtful according to
GBIF classification
Heteroprotaetia Mikšic 1963
Doubtful according to
GBIF classification
Heterorrhiniola Miksic 1974
Doubtful according to
GBIF classification
Heterosternus Dupont 1832
Doubtful according to
GBIF classification
Heterostetha Moser 1911
Doubtful according to
GBIF classification
Heterostheta
Doubtful according to
GBIF classification
Heterosyphus Paulian 1975
Doubtful according to
GBIF classification
Heteroychus
Doubtful according to
GBIF classification
Holcoribeus Nikritin 1977
Doubtful according to
GBIF classification
Holocnathon Martinez & Pereira 1956
Doubtful according to
GBIF classification
Holophylla Erichson 1847
Doubtful according to
GBIF classification
Holoschiza Lansberge 1886
Doubtful according to
GBIF classification
Homalothyrea Kolbe 1895
Doubtful according to
GBIF classification
Homioliogenys Gutierrez Alonso 1952
Doubtful according to
GBIF classification
Homocanthonides Luederwaldt 1929
Doubtful according to
GBIF classification
Homodesmius Sharp 1873
Doubtful according to
GBIF classification
Homoeopsopha Schürhoff 1934
Doubtful according to
GBIF classification
Homoliogenys Gutiérrez 1952
Doubtful according to
GBIF classification
Homolosternus Ohaus 1934
Doubtful according to
GBIF classification
Homosphileurus Kolbe 1910
Doubtful according to
GBIF classification
Hoplitogomphus Prell 1912
Doubtful according to
GBIF classification
Hoplocnemus Blanchard 1851
Doubtful according to
GBIF classification
Hoploscelis Burmeister 1844
Doubtful according to
GBIF classification
Hoplosternus Guerin-Meneville 1838
Doubtful according to
GBIF classification
Horonotus Burmeister 1847
Doubtful according to
GBIF classification
Horpnus Paulian 1951
Doubtful according to
GBIF classification
Hybalus Germar 1838
Doubtful according to
GBIF classification
Hybomidium Shipp 1897
Doubtful according to
GBIF classification
Hyperastia Fairmaire 1898
Doubtful according to
GBIF classification
Hypochrus Fairmaire 1891
Doubtful according to
GBIF classification
Hypopholis Erichson 1847
Doubtful according to
GBIF classification
Iapurea Martinez 1953
Doubtful according to
GBIF classification
Iarupoides Chalumeau 1984
Doubtful according to
GBIF classification
Idioschema Arrow 1914
Doubtful according to
GBIF classification
Ikelia Endrödi 1958
Doubtful according to
GBIF classification
Imelda Dellacasa, Gordon & Dellacasa 2002
Doubtful according to
GBIF classification
Incalidia Janson 1907
Doubtful according to
GBIF classification
Indocetonia Mikšic 1965
Doubtful according to
GBIF classification
Indoliocola Alexis & Delpont 1998
Doubtful according to
GBIF classification
Indoprotaetia
Doubtful according to
GBIF classification
Inhambane Péringuey 1907
Doubtful according to
GBIF classification
Insuloryctes Dechambre 2000
Doubtful according to
GBIF classification
Ipselissus
Doubtful according to
GBIF classification
Irrorhotides Shipp 1896
Doubtful according to
GBIF classification
Ischnestoma
Doubtful according to
GBIF classification
Ischnochelus Burmeister 1844
Doubtful according to
GBIF classification
Ischyrocera Hauser 1904
Doubtful according to
GBIF classification
Ishiopsopha
Doubtful according to
GBIF classification
Isodon Hope 1837
Doubtful according to
GBIF classification
Jansongora Paulian 1994
Doubtful according to
GBIF classification
Janssensantus Paulian 1976
Doubtful according to
GBIF classification
Jothochilus Kolbe 1892
Doubtful according to
GBIF classification
Krikkenica Delpont 1997
Doubtful according to
GBIF classification
Kuytenia Mikšic 1963
Doubtful according to
GBIF classification
Labathia Fairmaire 1898
Doubtful according to
GBIF classification
Lacris Fairmaire & Germain 1860
Doubtful according to
GBIF classification
Laonthophagus Frey 1971
Doubtful according to
GBIF classification
Latemnis Thomson 1880
Doubtful according to
GBIF classification
Latescutella Ruter 1972
Doubtful according to
GBIF classification
Laurentiana Ruter 1952
Doubtful according to
GBIF classification
Lecantotrogus Kolbe 1894
Doubtful according to
GBIF classification
Leptolepis Ley 1917
Doubtful according to
GBIF classification
Leptothyrea Kraatz 1882
Doubtful according to
GBIF classification
Leribe Péringuey 1904
Doubtful according to
GBIF classification
Lethosesthes Thomson 1880
Doubtful according to
GBIF classification
Leucochilus Kraatz 1897
Doubtful according to
GBIF classification
Leucoserica Reitter 1896
Doubtful according to
GBIF classification
Leuroserica Arrow 1947
Doubtful according to
GBIF classification
Liberocera Deloya & Ratcliffe 1988
Doubtful according to
GBIF classification
Lignarius Gistl 1848
Doubtful according to
GBIF classification
Ligyromorphus Arrow 1901
Doubtful according to
GBIF classification
Linotarsia Kraatz 1880
Doubtful according to
GBIF classification
Lipoclita Péringuey 1907
Doubtful according to
GBIF classification
Lipothyrea Péringuey 1907
Doubtful according to
GBIF classification
Liptothyrea
Doubtful according to
GBIF classification
Liteupatorus Prell 1911
Doubtful according to
GBIF classification
Loeblietus Endrodi 1979
Doubtful according to
GBIF classification
Lophogastra Fairmaire 1905
Doubtful according to
GBIF classification
Lophophora Kraatz 1895
Doubtful according to
GBIF classification
Lophorrhna Westwood 1842
Doubtful according to
GBIF classification
Luederwaldtina
Doubtful according to
GBIF classification
Luederwaldtinia Martínez 1951
Doubtful according to
GBIF classification
Luederwaltinia
Doubtful according to
GBIF classification
Macrocanius Schürhoff 1935
Doubtful according to
GBIF classification
Macrodactylus Berthold 1827
Doubtful according to
GBIF classification
Macroglycyphana Miksic 1968
Doubtful according to
GBIF classification
Macroliocola Alexis & Delpont 1998
Doubtful according to
GBIF classification
Macrominops Krikken
Doubtful according to
GBIF classification
Macrominus Mc Leay 1838
Doubtful according to
GBIF classification
Macronthophagus Ochi 2003
Doubtful according to
GBIF classification
Macropanelus Ochi, Kon & Araya 1998
Doubtful according to
GBIF classification
Macropocopris Arrow 1920
Doubtful according to
GBIF classification
Macropodina Casey 1915
Doubtful according to
GBIF classification
Macroprotaetia Mikšic 1965
Doubtful according to
GBIF classification
Macroryctes
Doubtful according to
GBIF classification
Macrotetrus Peringuey 1908
Doubtful according to
GBIF classification
Macrothops Dupont 1835
Doubtful according to
GBIF classification
Macrotrigonophorus Miksic 1974
Doubtful according to
GBIF classification
Madaphacosoma Paulian 1975
Doubtful according to
GBIF classification
Madateuchus Paulian 1953
Doubtful according to
GBIF classification
Magniscarabaeus Hong 1988
Doubtful according to
GBIF classification
Manodema Moser 1907
Doubtful according to
GBIF classification
Maraxes Lewis 1895
Doubtful according to
GBIF classification
Maripa Soula 2002
Doubtful according to
GBIF classification
Matashia Matsumura 1938
Doubtful according to
GBIF classification
Mazoe Péringuey 1896
Doubtful according to
GBIF classification
Mecaspidus Bourgoin 1921
Doubtful according to
GBIF classification
Mecopelidinota F. Bates 1904
Doubtful according to
GBIF classification
Mecynorrhinella Marais & Holm 1992
Doubtful according to
GBIF classification
Megalaemus Montrouzier 1860
Doubtful according to
GBIF classification
Megalleucosma Antoine 1989
Doubtful according to
GBIF classification
Megalorrhina
Doubtful according to
GBIF classification
Meganoxia Reitter 1902
Doubtful according to
GBIF classification
Meganthracophora
Doubtful according to
GBIF classification
Megaphanaeus d'Olsoufieff 1924
Doubtful according to
GBIF classification
Megaponerophilus Janssens 1949
Doubtful according to
GBIF classification
Megascarabaeus
Doubtful according to
GBIF classification
Megathopomima Martínez 1961
Doubtful according to
GBIF classification
Megathoposoma Balthasar 1939
Doubtful according to
GBIF classification
Melanochroea Kraatz 1900
Doubtful according to
GBIF classification
Melanomaladera Miyake & Yamaya 2001
Doubtful according to
GBIF classification
Melanoptera
Doubtful according to
GBIF classification
Melanosa Mulsant & Rey 1871
Doubtful according to
GBIF classification
Melichus Péringuey 1901
Doubtful according to
GBIF classification
Melinesthes Kraatz 1880
Doubtful according to
GBIF classification
Melolontha Müller 1764
Doubtful according to
GBIF classification
Meloroba
Doubtful according to
GBIF classification
Menthophilus
Doubtful according to
GBIF classification
Meretricula Özdikmen 2008
Doubtful according to
GBIF classification
Mesorhopa Kraatz 1880
Doubtful according to
GBIF classification
Mesorrhopa
Doubtful according to
GBIF classification
Metabolus
Doubtful according to
GBIF classification
Metaceraspis Frey 1962
Doubtful according to
GBIF classification
Metallophanaeus d'Olsoufieff 1924
Doubtful according to
GBIF classification
Mettalophanaeus
Doubtful according to
GBIF classification
Microcoenus Britton 1957
Doubtful according to
GBIF classification
Microcosmiomorpha Miksic 1974
Doubtful according to
GBIF classification
Microglycyphana Miksic 1968
Doubtful according to
GBIF classification
Micropeocila
Doubtful according to
GBIF classification
Microtertnitodius Pittino 2006
Doubtful according to
GBIF classification
Microthops
Doubtful according to
GBIF classification
Miksicus Özdikmen & Turgut 2009
Doubtful according to
GBIF classification
Millotsis Bourgin 1945
Doubtful according to
GBIF classification
Mimognorimus Kirkken
Doubtful according to
GBIF classification
Mimotribostethes Gutiérrez 1949
Doubtful according to
GBIF classification
Minilasiocala Soula 2006
Doubtful according to
GBIF classification
Minthophilus
Doubtful according to
GBIF classification
Miochaetopisthes Tangelder & Krikken 1982
Doubtful according to
GBIF classification
Mioxysternon Edmonds 1972
Doubtful according to
GBIF classification
Miridiba Coca-Abia 2008
Doubtful according to
GBIF classification
Mitrophorus Burmeister 1844
Doubtful according to
GBIF classification
Mixigenus Thomson 1859
Doubtful according to
GBIF classification
Moscheuma Thomson 1880
Doubtful according to
GBIF classification
Mucterophallus Poll 1886
Doubtful according to
GBIF classification
Mycterophalus
Doubtful according to
GBIF classification
Mycteroplus Fairmaire 1903
Doubtful according to
GBIF classification
Myctoplerus Fairmaire 1903
Doubtful according to
GBIF classification
Myodermidius Bourgoin 1921
Doubtful according to
GBIF classification
Myrmecotonus Mann 1914
Doubtful according to
GBIF classification
Nannovalgus Kolbe 1904
Doubtful according to
GBIF classification
Nataia Endrodi 1961
Doubtful according to
GBIF classification
Neateuchus Gillet 1911
Doubtful according to
GBIF classification
Neatocnemis Prell 1936
Doubtful according to
GBIF classification
Neoblitopertha Baraud 1991
Doubtful according to
GBIF classification
Neocavonus Blackburn 1888
Doubtful according to
GBIF classification
Neochaetopisthes Wasmann 1918
Doubtful according to
GBIF classification
Neocnemis Sharp 1875
Doubtful according to
GBIF classification
Neocymophorus Schein 1954
Doubtful according to
GBIF classification
Neodiapterina Dellacasa 1986
Doubtful according to
GBIF classification
Neodon Carne 1957
Doubtful according to
GBIF classification
Neomnematium Janssens 1938
Doubtful according to
GBIF classification
Neopachysoma Ferreira 1953
Doubtful according to
GBIF classification
Neopatatra Moore & Jameson 2013
Doubtful according to
GBIF classification
Neopentelia Matsumoto 2005
Doubtful according to
GBIF classification
Neophaedimus Schoch 1894
Doubtful according to
GBIF classification
Neopharis
Doubtful according to
GBIF classification
Neophoedimus Lucas 1870
Doubtful according to
GBIF classification
Neophonia Thomson 1878
Doubtful according to
GBIF classification
Neosaprosites Endrödi 1951
Doubtful according to
GBIF classification
Neosyrichthoschema Ferreira 1965
Doubtful according to
GBIF classification
Neotemnorrhynchus Janssens 1949
Doubtful according to
GBIF classification
Nephrodophus Sharp 1873
Doubtful according to
GBIF classification
Neprodopus
Doubtful according to
GBIF classification
Nesocanthon Pereira & Martínez 1956
Doubtful according to
GBIF classification
Nesopalla Paulian & Howden 1982
Doubtful according to
GBIF classification
Netocia Costa 1852
Doubtful according to
GBIF classification
Netociomima Mikšic 1963
Doubtful according to
GBIF classification
Niponoprotaetia Miksic 1966
Doubtful according to
GBIF classification
Notheterogomphus Prell 1912
Doubtful according to
GBIF classification
Notiophanaeus Edmonds 1994
Doubtful according to
GBIF classification
Notopedaria Matthews 1976
Doubtful according to
GBIF classification
Nselenius Holm & Perissinotto 2004
Doubtful according to
GBIF classification
Nudipleurus Lebis 1960
Doubtful according to
GBIF classification
Ochodaeus Serville 1825
Doubtful according to
GBIF classification
Ochranoxia Kraatz 1888
Doubtful according to
GBIF classification
Octodon Lansberge 1874
Doubtful according to
GBIF classification
Octotemna Blanchard 1850
Doubtful according to
GBIF classification
Odontoprotaetia Mikšic 1963
Doubtful according to
GBIF classification
Odontothyrea Schoch 1897
Doubtful according to
GBIF classification
Oedanomerus Sehnal 2018
Doubtful according to
GBIF classification
Okeurophorus
Doubtful according to
GBIF classification
Oniticellus Le Peletier & Serville 1828
Doubtful according to
GBIF classification
Onochirus Kolbe 1891
Doubtful according to
GBIF classification
Onthohpagus
Doubtful according to
GBIF classification
Onthophagus
Doubtful according to
GBIF classification
,
GBIF classification
,
GBIF classification
,
GBIF classification
,
GBIF classification
,
GBIF classification
,
GBIF classification
,
GBIF classification
, and
GBIF classification
Onthophahus
Doubtful according to
GBIF classification
Onychotarsus Schoch 1897
Doubtful according to
GBIF classification
Oogeneius Solier 1851
Doubtful according to
GBIF classification
Opistocus Britton 1957
Doubtful according to
GBIF classification
Orchiserica Miyake & Yamaya 2001
Doubtful according to
GBIF classification
Oreopotosia Medvedev 1964
Doubtful according to
GBIF classification
Orphnidius Benderitter 1913
Doubtful according to
GBIF classification
Ortheterogomphus Prell 1912
Doubtful according to
GBIF classification
Oruscatuen Bates 1870
Doubtful according to
GBIF classification
Oryctes Fabricius
Doubtful according to
GBIF classification
Oryctus
Doubtful according to
GBIF classification
Otoclinus Brenske 1896
Doubtful according to
GBIF classification
Oxyomus Kirby & Spence 1828
Doubtful according to
GBIF classification
Oxyperatex Krikken 1982
Doubtful according to
GBIF classification
Oxyrraphia
Doubtful according to
GBIF classification
Pachnodella Stichel 1922
Doubtful according to
GBIF classification
Pachnotosia Reitter 1891
Doubtful according to
GBIF classification
Pachycerus Guérin
Doubtful according to
GBIF classification
Pachylomerus Bertolini 1849
Doubtful according to
GBIF classification
Pachylotoma Blanchard 1850
Doubtful according to
GBIF classification
Pachyorphnus Paulian 1951
Doubtful according to
GBIF classification
Pachyprotaetia Mikšic 1965
Doubtful according to
GBIF classification
Paediovalgus Kolbe 1904
Doubtful according to
GBIF classification
Pagonotarsus Burmeister 1842
Doubtful according to
GBIF classification
Pahangia Arrow 1948
Doubtful according to
GBIF classification
Paidiovalgus
Doubtful according to
GBIF classification
Palaeonthophagus Zunino 1979
Doubtful according to
GBIF classification
Palmerstonia Blackburn 1888
Doubtful according to
GBIF classification
Parabothynus Endrodi 1968
Doubtful according to
GBIF classification
Paradendropaemon Edmonds 1972
Doubtful according to
GBIF classification
Paragynaecoserica Khan & Ghai 1982
Doubtful according to
GBIF classification
Parahyboma Paulian 1938
Doubtful according to
GBIF classification
Paraliatongus Balthasar 1963
Doubtful according to
GBIF classification
Parapanelus Balthasar 1966
Doubtful according to
GBIF classification
Parapeltastes Ricchiardi, Perissinotto & Clennell 2004
Doubtful according to
GBIF classification
Paraphanaemorphus
Doubtful according to
GBIF classification
Parapinotus Harold 1878
Doubtful according to
GBIF classification
Parapsammodius Verdú, Stebnicka & Galante 2006
Doubtful according to
GBIF classification
Parascarabaeus Balthasar 1961
Doubtful according to
GBIF classification
Paraspidolea Höhne 1922
Doubtful according to
GBIF classification
Parbomonyx Ohaus 1915
Doubtful according to
GBIF classification
Parentius Zunino 1979
Doubtful according to
GBIF classification
Pareurysternus Martinez 1988
Doubtful according to
GBIF classification
Parodochilus Rakovic 1997
Doubtful according to
GBIF classification
Paronitis Balthasar 1942
Doubtful according to
GBIF classification
Paronthophagus Balthasar 1959
Doubtful according to
GBIF classification
Parorphnus Paulian 1951
Doubtful according to
GBIF classification
Parvulomaladera Ahrens 1995
Doubtful according to
GBIF classification
Passalaphodius Martínez 1953
Doubtful according to
GBIF classification
Peltecanthon Pereira 1953
Doubtful according to
GBIF classification
Peltotrichius
Doubtful according to
GBIF classification
Peltotrichus Howden 1968
Doubtful according to
GBIF classification
Penrithochilus Krikken 1990
Doubtful according to
GBIF classification
Peradoretus Reitter 1903
Doubtful according to
GBIF classification
Petroyitzia Miksic 1965
Doubtful according to
GBIF classification
Peyrierasantus Paulian 1976
Doubtful according to
GBIF classification
Phacosomoides Balthasar 1966
Doubtful according to
GBIF classification
Phacosomoides Martínez & Pereira 1959
Doubtful according to
GBIF classification
Phaedotrodus
Doubtful according to
GBIF classification
Phaedotrogus Paulian 1985
Doubtful according to
GBIF classification
Phanaresthes
Doubtful according to
GBIF classification
Philochlaenea Hope 1839
Doubtful according to
GBIF classification
Phornus Paulian 1951
Doubtful according to
GBIF classification
Phyllopodium Schoch 1895
Doubtful according to
GBIF classification
Phylloputha Kirby & Spence 1828
Doubtful according to
GBIF classification
Phymatopterix
Doubtful according to
GBIF classification
Pilleriana Baraud 1991
Doubtful according to
GBIF classification
Plaesiorrhina Westwood 1842
Doubtful according to
GBIF classification
Plaesiorrhinella
Doubtful according to
GBIF classification
Planontherus Genier 1996
Doubtful according to
GBIF classification
Platedelosis Kraatz 1880
Doubtful according to
GBIF classification
Platyphymatica
Doubtful according to
GBIF classification
Plesioserica Brenske 1900
Doubtful according to
GBIF classification
Plinurgus
Doubtful according to
GBIF classification
Plocovalgus Kolbe 1904
Doubtful according to
GBIF classification
Podoserica Breit 1912
Doubtful according to
GBIF classification
Poecilophana Kraatz 1895
Doubtful according to
GBIF classification
Poecilophila Kolbe 1893
Doubtful according to
GBIF classification
Poecilophilides Kraatz 1898
Doubtful according to
GBIF classification
Polybaphes Kirby 1827
Doubtful according to
GBIF classification
Polybathes Kirby 1827
Doubtful according to
GBIF classification
Polylamina Hardy 1974
Doubtful according to
GBIF classification
Polystigma Kraatz 1880
Doubtful according to
GBIF classification
Ponerotrogus Silvestri 1924
Doubtful according to
GBIF classification
Porphyrobapta Kraatz 1888
Doubtful according to
GBIF classification
Potosiomima
Doubtful according to
GBIF classification
Praelinotarsia Duvivier 1891
Doubtful according to
GBIF classification
Praogolofa Bates 1892
Doubtful according to
GBIF classification
Pristadoretus Reitter 1903
Doubtful according to
GBIF classification
Progastor Thomson 1880
Doubtful according to
GBIF classification
Propomacus
Doubtful according to
GBIF classification
Protaetiola Mikšic 1963
Doubtful according to
GBIF classification
Protaetiophana Heller 1896
Doubtful according to
GBIF classification
Proteroscarabeus Grabau 1923
Doubtful according to
GBIF classification
Protoctenius Medvedev 1952
Doubtful according to
GBIF classification
Protoleura Britton 1987
Doubtful according to
GBIF classification
Pseudagoliinus
Doubtful according to
GBIF classification
Pseudanatona Kraatz 1895
Doubtful according to
GBIF classification
Pseudanthracophora Kraatz 1898
Doubtful according to
GBIF classification
Pseudaplasta Kraatz 1898
Doubtful according to
GBIF classification
Pseudarachnodes Lebis 1953
Doubtful according to
GBIF classification
Pseudastoxenus Bourgoin 1921
Doubtful according to
GBIF classification
Pseudepilissus
Doubtful according to
GBIF classification
Pseudocalopotosia Miksic 1966
Doubtful according to
GBIF classification
Pseudocetonischema Mikšic 1965
Doubtful according to
GBIF classification
Pseudochironitis Ferreira 1977
Doubtful according to
GBIF classification
Pseudocoptorhina Ferreira 1954
Doubtful according to
GBIF classification
Pseudocymophorus Schein 1954
Doubtful according to
GBIF classification
Pseudocyphonistes Péringuey 1901
Doubtful according to
GBIF classification
Pseudodontoplia Lebis 1961
Doubtful according to
GBIF classification
Pseudoeuselates Miksic 1974
Doubtful according to
GBIF classification
Pseudoheliocopris Ferreira 1970
Doubtful according to
GBIF classification
Pseudohercitis Moser 1921
Doubtful according to
GBIF classification
Pseudoheterophana Allard 1990
Doubtful according to
GBIF classification
Pseudoholophylla Blackburn 1911
Doubtful according to
GBIF classification
Pseudoisonychus Frey 1971
Doubtful according to
GBIF classification
Pseudoleuretra Martínez & d'Andretta 1955
Doubtful according to
GBIF classification
Pseudolontha Fairmaire 1897
Doubtful according to
GBIF classification
Pseudoneptunides Csiki 1909
Doubtful according to
GBIF classification
Pseudonetocia Medvedev 1964
Doubtful according to
GBIF classification
Pseudonthophagus Balthasar 1959
Doubtful according to
GBIF classification
Pseudopantrogus Petrovitz 1969
Doubtful according to
GBIF classification
Pseudophacosoma
Doubtful according to
GBIF classification
Pseudoprotaetia Kraatz 1882
Doubtful according to
GBIF classification
Pseudopygora Kraatz 1897
Doubtful according to
GBIF classification
Pseudorhomborrhina Miksic 1967
Doubtful according to
GBIF classification
Pseudorycytes
Doubtful according to
GBIF classification
Pseudosaprosites Balthasar 1972
Doubtful according to
GBIF classification
Pseudourbania Mikšic 1965
Doubtful according to
GBIF classification
Psileterogomphus Prell 1912
Doubtful according to
GBIF classification
Psuedotephraea Kraatz 1882
Doubtful according to
GBIF classification
Pteronyx
Doubtful according to
GBIF classification
Pterothochaetes
Doubtful according to
GBIF classification
Ptychodestes
Doubtful according to
GBIF classification
Ptychophorus Burmeister & Schaum 1841
Doubtful according to
GBIF classification
Puapana Heller 1913
Doubtful according to
GBIF classification
Pulgilotrichius Krikken
Doubtful according to
GBIF classification
Pygoropsis Schoch 1896
Doubtful according to
GBIF classification
Pyropotosia Reitter 1898
Doubtful according to
GBIF classification
Relictonthophagus Kabakov 1979
Doubtful according to
GBIF classification
Reneria
Doubtful according to
GBIF classification
Rhinaspoides Moser 1919
Doubtful according to
GBIF classification
Rhomborrhine
Doubtful according to
GBIF classification
Rhyncoserica Burgeon 1942
Doubtful according to
GBIF classification
Rhyncosymmela Frey 1974
Doubtful according to
GBIF classification
Robinsonella Lacroix 1993
Doubtful according to
GBIF classification
Ronphus Paulian 1951
Doubtful according to
GBIF classification
Rubrohyboma Paulian 1939
Doubtful according to
GBIF classification
Rutelorcytes Arrow 1908
Doubtful according to
GBIF classification
Ruteraetia Krikken 1980
Doubtful according to
GBIF classification
Rykanes Minck 1919
Doubtful according to
GBIF classification
Rykanoryctes Minck 1919
Doubtful according to
GBIF classification
Ryteloryctes
Doubtful according to
GBIF classification
Saproecius Péringuey 1901
Doubtful according to
GBIF classification
Saprosotellus Balthasar 1967
Doubtful according to
GBIF classification
Saprosties Redtenbacher 1858
Doubtful according to
GBIF classification
Sarothermus Blackburn 1907
Doubtful according to
GBIF classification
Scaptophilus Burmeister 1847
Doubtful according to
GBIF classification
Scarabaeidae
Harvest ancestor according to
Global Invasive Species Database
,
Arkive
,
EOL authors
,
EOL authors
,
Animal Diversity Web
,
Global Register of Introduced and Invasive Species
,
GBIF national node type records UK
,
GBIF national node type records France
,
GBIF national node type records Brazil
,
GBIF national node type records Sweden
,
Mozambique Species List
,
GBIF nat'l node type records Netherlands
,
GBIF national node type records Germany
,
NMNH type specimens
,
IUCN
,
Zookeys
,
Zookeys
,
iDigBio type specimen records
,
iDigBio type specimen records
,
iDigBio type specimen records
,
iDigBio type specimen records
,
Paleobiology Database
,
CITES
,
Animal Diversity Web Traits and images
,
iNaturalist
,
NMNH Collection
,
Harvard Museum of Comparative Zoology
,
Alaskan Arthropods
,
Simpson et al 2020
,
BioImages, the virtual fieldguide, UK
,
TreatmentBank
,
Global Biotic Interactions
,
Global Biotic Interactions
,
Global Biotic Interactions
, and
Flickr Group
Scarabaeidae Latreille 1802
Harvest ancestor according to
EOL authors
and
EOL authors
Scarabaes
Doubtful according to
GBIF classification
Schizonycha Blanchard 1845
Doubtful according to
GBIF classification
Schizorrhina Burmeister 1842
Doubtful according to
GBIF classification
Schoenherria Burmeister 1855
Doubtful according to
GBIF classification
Sebasteos Westwood 1845
Doubtful according to
GBIF classification
Selenocopris Burmeister 1846
Doubtful according to
GBIF classification
Sericaria Motschoulsky 1860
Doubtful according to
GBIF classification
Seudinca
Doubtful according to
GBIF classification
Shangaia Lucas 1920
Doubtful according to
GBIF classification
Sikorantus Paulian 1976
Doubtful according to
GBIF classification
Simogonious Harold 1871
Doubtful according to
GBIF classification
Singhalopotosia Miksic 1987
Doubtful according to
GBIF classification
Sinoliocola Alexis & Delpont 1998
Doubtful according to
GBIF classification
Sinoplia Medvedev 1952
Doubtful according to
GBIF classification
Sinoserica Miyake & Yamaya 2001
Doubtful according to
GBIF classification
Sissantobius Ritsema 1888
Doubtful according to
GBIF classification
Sisyphe Latreille 1807
Doubtful according to
GBIF classification
Sisyphus Wiedemann 1823
Doubtful according to
GBIF classification
Sisyraphora Kraatz 1881
Doubtful according to
GBIF classification
Spaeromorphus
Doubtful according to
GBIF classification
Spermohybosorus Pic 1922
Doubtful according to
GBIF classification
Sphaerocanthon
Doubtful according to
GBIF classification
Sphaeromrophus
Doubtful according to
GBIF classification
Spherocanthon Olsoufieff 1947
Doubtful according to
GBIF classification
Sphyrorrhina Nickerl 1891
Doubtful according to
GBIF classification
Spilovalgus Kolbe 1904
Doubtful according to
GBIF classification
Spinohoplia Sabatinelli 1997
Doubtful according to
GBIF classification
Stenocorythoderus Wasmann 1918
Doubtful according to
GBIF classification
Stenolomaptera Kraatz 1898
Doubtful according to
GBIF classification
Stephanocrates Kolbe 1892
Doubtful according to
GBIF classification
Stephopseudinca Lucassen 1933
Doubtful according to
GBIF classification
Sternoplidius Moser 1908
Doubtful according to
GBIF classification
Stichadoretus Reitter 1903
Doubtful according to
GBIF classification
Stichothyrea Kraatz 1882
Doubtful according to
GBIF classification
Stizopygora Kraatz 1881
Doubtful according to
GBIF classification
Strategus Hope 1837
Doubtful according to
GBIF classification
Streblopoides Balthasar 1938
Doubtful according to
GBIF classification
Strehlia Frey 1969
Doubtful according to
GBIF classification
Strongylothorax Motschoulsky 1860
Doubtful according to
GBIF classification
Stypotrupes Burmeister 1847
Doubtful according to
GBIF classification
Subpeltonotus Swaraj Ghai, Kailash Chandra & Ramamurthy 1988
Doubtful according to
GBIF classification
Sulcodrepanus Krikken 2009
Doubtful according to
GBIF classification
Sunenaga Ochi 2003
Doubtful according to
GBIF classification
Surutoides Endrodi 1981
Doubtful according to
GBIF classification
Svobia Mikšic 1965
Doubtful according to
GBIF classification
Syneterogomphu
Doubtful according to
GBIF classification
Syntomopteryx Kraatz 1900
Doubtful according to
GBIF classification
Syrichtomorphus
Doubtful according to
GBIF classification
Syrichtus Kirby 1837
Doubtful according to
GBIF classification
Syrictoides Endrodi 1977
Doubtful according to
GBIF classification
Systellorhina Kraatz 1895
Doubtful according to
GBIF classification
Taenia Stebnicka 2003
Doubtful according to
GBIF classification
Taeniesthes Kraatz 1880
Doubtful according to
GBIF classification
Tamisoria Thomson 1877
Doubtful according to
GBIF classification
Tanzanolus Scholtz & Howden 1987
Doubtful according to
GBIF classification
Taphraeoserica Brenske 1900
Doubtful according to
GBIF classification
Taurocopris d'Olsoufieff 1924
Doubtful according to
GBIF classification
Taurrhina
Doubtful according to
GBIF classification
Taxipertha Baraud 1991
Doubtful according to
GBIF classification
Telhyboma Kolbe 1893
Doubtful according to
GBIF classification
Temnorrhynchodes Prell 1934
Doubtful according to
GBIF classification
Temnostoma Blanchard 1850
Doubtful according to
GBIF classification
Teraserica Brenske 1899
Doubtful according to
GBIF classification
Termitotrox Wasmann 1918
Doubtful according to
GBIF classification
Tetraaechma Blanchard 1846
Doubtful according to
GBIF classification
Tetraechma Agassiz 1845
Doubtful according to
GBIF classification
Tetragonorhina Kraatz 1896
Doubtful according to
GBIF classification
Tetrarhabdotis Kraatz 1883
Doubtful according to
GBIF classification
Tetrodontus Zhang 1991
Doubtful according to
GBIF classification
Tetrophthalmus Kirby 1827
Doubtful according to
GBIF classification
Thaiaphodius Masumoto 1991
Doubtful according to
GBIF classification
Thamnastopeos Kraatz 1883
Doubtful according to
GBIF classification
Tibiovalgus Krikken 1978
Doubtful according to
GBIF classification
Tichocelis Moser 1908
Doubtful according to
GBIF classification
Tomentoprotaetia Miksic 1987
Doubtful according to
GBIF classification
Tracheterogomphus Prell 1912
Doubtful according to
GBIF classification
Trachychlaenia Kraatz 1895
Doubtful according to
GBIF classification
Trapezorrhina Hauser 1904
Doubtful according to
GBIF classification
Triaphodius
Doubtful according to
GBIF classification
Tribostethes Curtis 1844
Doubtful according to
GBIF classification
Trichocanthon Pereira & Martinez 1960
Doubtful according to
GBIF classification
Trichocelis Moser 1908
Doubtful according to
GBIF classification
Trichomia
Doubtful according to
GBIF classification
Trichonthophagus Zunino 1979
Doubtful according to
GBIF classification
Trichoserica Reitter 1896
Doubtful according to
GBIF classification
Trichotarsia Burmeister 1842
Doubtful according to
GBIF classification
Trichothyrea Kolbe 1895
Doubtful according to
GBIF classification
Triclirea Schoch 1896
Doubtful according to
GBIF classification
Trigonochilus Brenske 1896
Doubtful according to
GBIF classification
Trigonocnemus Kraatz 1894
Doubtful according to
GBIF classification
Trionychus Burmeister 1847
Doubtful according to
GBIF classification
Trissodon Prell 1936
Doubtful according to
GBIF classification
Trochaloserica Brenske 1900
Doubtful according to
GBIF classification
Tropicalia Koçak & Kemal 2008
Doubtful according to
GBIF classification
Tropidonitis Janssens 1937
Doubtful according to
GBIF classification
Tryssus Erichson 1847
Doubtful according to
GBIF classification
Ulomenes Blanchard 1850
Doubtful according to
GBIF classification
Ulopterus Burmeister 1842
Doubtful according to
GBIF classification
Unguiculenaris Dewailly 1950
Doubtful according to
GBIF classification
Urbania Mikšic 1963
Doubtful according to
GBIF classification
Vansonia Schein 1956
Doubtful according to
GBIF classification
Vertumnus Reiche 1859
Doubtful according to
GBIF classification
Vietnamoprotaetia Mikšic 1971
Doubtful according to
GBIF classification
Vittighana Endrodi 1973
Doubtful according to
GBIF classification
Vulcanoia Martinez 1954
Doubtful according to
GBIF classification
Watkinsia Covington 1994
Doubtful according to
GBIF classification
Wernerophylla Lacroix 2001
Doubtful according to
GBIF classification
Widagdous Alexis & Delpont 1998
Doubtful according to
GBIF classification
Xenogenius Kolbe 1892
Doubtful according to
GBIF classification
Xiphoscelidus Péringuey 1907
Doubtful according to
GBIF classification
Xylorectes
Doubtful according to
GBIF classification
Xynedria Blackburn 1894
Doubtful according to
GBIF classification
Xynophron Harold 1880
Doubtful according to
GBIF classification
Zabacana Saylor 1946
Doubtful according to
GBIF classification
Zanitanus Lacroix 2001
Doubtful according to
GBIF classification
Zhangia Bunalski 2002
Doubtful according to
GBIF classification
Common Names
English
scarab beetles
preferred
Recognized by
English Vernaculars for Landmark Taxa
Arabic
جعليات
preferred
Recognized by
Wikidata Common Names
Armenian
թերթաբեղիկավորներ
preferred
Recognized by
Wikidata Common Names
Azerbaijani
Lövhəbığ böcəklər
preferred
Recognized by
Wikidata Common Names
Belarusian
Пласцініставусыя
preferred
Recognized by
Wikidata Common Names
пласткавусыя
prefer
Recognized by
Wikidata Common Names (prefers this name)
Bulgarian
Листороги бръмбари
preferred
Recognized by
Wikidata Common Names
Catalan; Valencian
escarabeids
preferred
Recognized by
Wikidata Common Names
Chinese
金龟科
preferred
Recognized by
Wikidata Common Names
Croatian
Truležari
preferred
Recognized by
Wikidata Common Names
Czech
vrubounovití
preferred
Recognized by
Wikidata Common Names
Danish
Torbister
preferred
Recognized by
Wikidata Common Names
Dinka
Aluotweer
preferred
Recognized by
Wikidata Common Names
Dutch; Flemish
Bladsprietkevers
preferred
Recognized by
Wikidata Common Names
Esperanto
Skarabedoj
preferred
Recognized by
Wikidata Common Names
Estonian
Põrniklased
preferred
Recognized by
Wikidata Common Names
Finnish
Lehtisarviset
preferred
Recognized by
Wikidata Common Names
and
admin
Galician
Escarabeidos
preferred
Recognized by
Wikidata Common Names
Georgian
ფირფიტულვაშა ხოჭოსებრნი
preferred
Recognized by
Wikidata Common Names
German
Blatthornkäfer
preferred
Recognized by
Wikidata Common Names
Hebrew
זבליתיים
preferred
Recognized by
Wikidata Common Names
Hungarian
ganajtúrófélék
preferred
Recognized by
Wikidata Common Names
Ido
Skarabeo
preferred
Recognized by
Wikidata Common Names
Irish
deá
preferred
Recognized by
Wikidata Common Names
Japanese
コガネムシ科
preferred
Recognized by
Furui 2020
,
Wikidata Common Names
, and
admin
Korean
풍뎅이과
preferred
Recognized by
Wikidata Common Names
Latvian
Skarabeju dzimta
preferred
Recognized by
Wikidata Common Names
Lithuanian
Skarabėjai
preferred
Recognized by
Wikidata Common Names
Low Saxon
Scarabäuskävers
preferred
Recognized by
Wikidata Common Names
Macedonian
тркалачи
preferred
Recognized by
Wikidata Common Names
Navajo; Navaho
Chéłchaaʼ bideeʼ nteelígíí
preferred
Recognized by
Wikidata Common Names
Norwegian
skarabidar
preferred
Recognized by
Wikidata Common Names
skarabider
preferred
Recognized by
Wikidata Common Names
Ossetian; Ossetic
Цъарæгрихионтæ
preferred
Recognized by
Wikidata Common Names
Persian
سرگینچرخانان
preferred
Recognized by
Wikidata Common Names
Portuguese
Escaravelho
preferred
Recognized by
Wikidata Common Names
Quechua
Raphiwaqracha suntu
preferred
Recognized by
Wikidata Common Names
Russian
пластинчатоусые
preferred
Recognized by
Wikidata Common Names
Serbian
Балегари
preferred
Recognized by
Wikidata Common Names
Slovak
Skarabeusovité
preferred
Recognized by
Wikidata Common Names
Spanish; Castilian
Skarabeji
preferred
Recognized by
Wikidata Common Names
Swedish
Bladhorningar
preferred
Recognized by
Wikidata Common Names
Ukrainian
Пластинчастовусі
preferred
Recognized by
Wikidata Common Names
Vietnamese
Họ Bọ hung
preferred
Recognized by
Wikidata Common Names
Wu
金龟子
preferred
Recognized by
Wikidata Common Names
Zhuang; Chuang
Duzmboenj
preferred
Recognized by
Wikidata Common Names
Arz
جعليات
preferred
Recognized by
Wikidata Common Names
Lzh
金龜
preferred
Recognized by
Wikidata Common Names
Mzn
گوزنگو
preferred
Recognized by
Wikidata Common Names
Nan
Kim-ku
preferred
Recognized by
Wikidata Common Names
Yue
金甲蟲
preferred
Recognized by
Wikidata Common Names
Zh Cn
金龟科
preferred
Recognized by
Wikidata Common Names
Zh Tw
金龜科
preferred
Recognized by
Wikidata Common Names
show 48 more languages
hide 48 languages
Curated hierarchies for Scarabaeidae
EOL Dynamic Hierarchy April 2022
Life
Cellular Organisms
Eukaryota
Opisthokonta
Metazoa
Bilateria
Protostomia
Ecdysozoa
Arthropoda
Pancrustacea
Hexapoda
Insecta
Pterygota
Neoptera
Endopterygota
Coleoptera
Polyphaga
Scarabaeiformia
Scarabaeoidea
Scarabaeidae
(this page)
Acanthonitis
Afrodrepanus
Agamopus
Aliuscanthoniola
Allogymnopleurus
Allonitis
Alloscelus
Amietina
Amphiceratodon
Amphistomus
Anisocanthon
Anoctus
Anomiopsoides
Anomiopus
Anonychonitis
Anoplodrepanus
Aphengium
Aphengoecus
Apotolamprus
Aptenocanthon
Apterepilissus
Aptychonitis
Arachnodes
Ateuchetus
Ateuchus
Attavicinus
Aulacopris
Ausmontins
Baloghonthobium
Bdelyropsis
Bdelyrus
Besourenga
Bohepilissus
Bolbites
Boletoscapter
Bradypodidium
Bubas
Byrrhidium
Caccobiomorphus
Caccobius
Caeconthobium
Cambefortantus
Cambefortius
Canthidium
Canthochilum
Canthodimorpha
Canthon
Canthonella
Canthonidia
Canthonosoma
Canthotrypes
Catharsiocopris
Catharsius
Cephalodesmius
Chalcocopris
Chalconotus
Cheironitis
Chiron
Circellium
Cleptocaccobius
Clypeodrepanus
Copridaspidus
Copris
Coproecus
Coprophanaeus
Coptodactyla
Coptorhina
Cryptocanthon
Cyobius
Cyptochirus
Degallieridium
Delopleurus
Deltepilissus
Deltochilum
Deltorhinum
Demarziella
Dendropaemon
Diabroctis
Diastellopalpus
Dichotomius
Dicranocara
Digitonthophagus
Diorygopyx
Disphysema
Dorbignyolus
Drepanocerus
Drepanoplatynus
Drepanopodus
Drogo
Dwesasilvasedis
Endroedyolus
Ennearabdus
Eodrepanus
Epactoides
Epidrepanus
Epilissus
Epirinus
Escarabaeus
Eucranium
Eudinopus
Euoniticellus
Euonthophagus
Eurysternus
Eusaproecius
Eutrichillum
Falsignambia
Feeridium
Frankenbergerius
Garreta
Genieridium
Gilletellus
Glyphoderus
Gromphas
Gymnopleurus
Gyronotus
Hammondantus
Hamonthophagus
Hansreia
Haroldius
Helictopleurus
Heliocopris
Heteroclitopus
Heteronitis
Holocanthon
Holocephalus
Homalotarsus
Homocopris
Homocorpis
Hyalonthophagus
Hypocanthidium
Ignambia
Isocopris
Ixodina
Janssensantus
Janssensellus
Jossonthophagus
Kheper
Kolbeellus
Krikkenius
Kurtops
Labroma
Larhodius
Latodrepanus
Leotrichillum
Lepanus
Liatongus
Litocopris
Lophodonitis
Macroderes
Malagoniella
Martinezidium
Matthewsius
Megalonitis
Megatharsis
Megathopa
Megathoposoma
Melanocanthon
Mentophilus
Metacatharsius
Microcopris
Milichus
Mimonthophagus
Mnematidium
Mnematium
Monoplistes
Monteithocanthon
Morettius
Namakwanus
Namaphilus
Nanos
Nebulasilvius
Neonitis
Neosaproecius
Neosisyphus
Nesosisyphus
Nesovinsonia
Nitiocellus
Nunoidium
Ochicanthon
Odontoloma
Oficanthon
Oniticellus
Onitis
Onoreidium
Ontherus
Onthobium
Onthophagus
Onychothecus
Oruscatus
Outenikwanus
Oxysternon
Pachylomera
Pachysoma
Panelus
Paracanthon
Parachorius
Paracryptocanthon
Paragymnopleurus
Paraixodina
Paraphacosomoides
Paraphytus
Parascatonomus
Parateuchus
Paroniticellus
Paronthobium
Parvuhowdenius
Peckolus
Pedaria
Pedaridium
Penalus
Pereiraidium
Phaedotrogus
Phalops
Phanaeus
Pinacopodius
Pinacotarsus
Platyonitis
Pleuronitis
Proagoderus
Pseudignambia
Pseudocanthon
Pseudochironitis
Pseudocopris
Pseudonthobium
Pseudopedaria
Pseudosaproecius
Pseuduroxys
Pycnopanelus
Saphobiamorpha
Saphobius
Sarophorus
Sauvagesinella
Scaptocnemis
Scaptodera
Scarabaeolus
Scarabaeus
Scatimus
Scatonomus
Scatrichus
Sceliages
Scybalocanthon
Scybalophagus
Silvaphilus
Silvinha
Sinapisoma
Sinodrepanus
Sisyphus
Stiptocnemis
Stiptopodius
Stiptotarsus
Streblopus
Sukelus
Sulcophanaeus
Sylvicanthon
Synapsis
Tanzanolus
Temnoplectron
Tesserodon
Tesserodoniella
Tetraechma
Theotimius
Thyregis
Tibiodrepanus
Tiniocellus
Tomogonus
Tragiscus
Trichillidium
Trichillum
Tropidonitis
Uenoeus
Unidentis
Uroxys
Versicorpus
Vulcanocanthon
Walterantus
Xenocanthon
Xinidium
Yvescambefortius
Zonocopris
Aclopidae
Aegialiidae
Alloioscarabaeidae
Aphodiidae
Aulonocnemidae
Belohinidae
Bolboceratidae
Cetoniidae
Diphyllostomatidae
Dynastidae
Eremazidae
Euchiridae
Geotrupidae
Glaphyridae
Glaresidae
Hybosoridae
Lucanidae
Melolonthidae
Ochodaeidae
Orphnidae
Pachypodidae
Passalidae
Passalopalpidae
Pleocomidae
Rutelidae
Septiventeridae
Termitotrogidae
Trogidae
Catalogue of Life 26 March 2024
Biota
Animalia
Arthropoda
Insecta
Coleoptera
Scarabaeoidea
Scarabaeidae Latreille 1802
(this page)
Aclopinae Blanchard 1850
Aegialiinae Castelnau 1840
Allidiostomatinae Arrow 1940
Antiqusolidus Lu, Ahrens, Bai, Shih & Ren 2023
Aphodiinae Leach 1815
Aulonocneminae Janssens 1946
Cetoniinae Leach 1815
Chironinae Blanchard 1845
Cretoscarabaeinae Nikolajev 1995
Dynamopodinae Arrow 1911
Dynastinae MacLeay 1819
Eremazinae Stebnicka 1977
Hopliinae Latreille 1829
Hybosorites Nikolajev 1996
Lithoscarabaeinae Nikolajev 1992
Melolonthinae MacLeay 1819
Oncerinae Le Conte 1861
Orphninae Erichson 1847
Pachydeminae Reitter 1902
Phaenomeridinae Erichson 1847
Podolasiinae Howden 1997
Prototroginae Nikolajev 2000
Rutelinae MacLeay 1819
Scarabaeinae Latreille 1802
Termitotroginae Wasmann 1918
Trichiinae Kolbe 1897
Valginae Schenkling 1922
Alloioscarabaeidae Bai, Ren & Yang 2012
Belohinidae Paulian 1958
Diphyllostomatidae Holloway 1972
Geotrupidae Latreille 1802
Glaphyridae MacLeay 1819
Glaresidae Preudhomme de Borre 1886
Hybosoridae Erichson 1847
Lucanidae Latreille 1804
Ochodaeidae Streubel 1846
Passalidae Leach 1815
Passalopalpidae Boucher & Bai 2016
Pleocomidae Le Conte 1861
Septiventeridae Bai, Ren, Shih & Yang 2013
Trogidae MacLeay 1819
Dynamic Hierarchy Version 2.2.3
Scarabaeidae
(this page)
Acanthonitis
Afrodrepanus
Agamopus
Aliuscanthoniola
Allogymnopleurus
Allonitis
Alloscelus
Amietina
Amphiceratodon
Amphistomus
Anisocanthon
Anoctus
Anomiopsoides
Anomiopus
Anonychonitis
Anoplodrepanus
Aphengium
Aphengoecus
Apotolamprus
Aptenocanthon
Apterepilissus
Aptychonitis
Arachnodes
Ateuchetus
Ateuchus
Attavicinus
Aulacopris
Ausmontins
Baloghonthobium
Bdelyropsis
Bdelyrus
Besourenga
Bohepilissus
Bolbites
Boletoscapter
Bradypodidium
Bubas
Byrrhidium
Caccobiomorphus
Caccobius
Caeconthobium
Cambefortantus
Cambefortius
Canthidium
Canthochilum
Canthodimorpha
Canthon
Canthonella
Canthonidia
Canthonosoma
Canthotrypes
Catharsiocopris
Catharsius
Cephalodesmius
Chalcocopris
Chalconotus
Cheironitis
Chiron
Circellium
Cleptocaccobius
Clypeodrepanus
Copridaspidus
Copris
Coproecus
Coprophanaeus
Coptodactyla
Coptorhina
Cryptocanthon
Cyobius
Cyptochirus
Degallieridium
Delopleurus
Deltepilissus
Deltochilum
Deltorhinum
Demarziella
Dendropaemon
Diabroctis
Diastellopalpus
Dichotomius
Dicranocara
Digitonthophagus
Diorygopyx
Disphysema
Dorbignyolus
Drepanocerus
Drepanoplatynus
Drepanopodus
Drogo
Dwesasilvasedis
Endroedyolus
Ennearabdus
Eodrepanus
Epactoides
Epidrepanus
Epilissus
Epirinus
Escarabaeus
Eucranium
Eudinopus
Euoniticellus
Euonthophagus
Eurysternus
Eusaproecius
Eutrichillum
Falsignambia
Feeridium
Frankenbergerius
Garreta
Genieridium
Gilletellus
Glyphoderus
Gromphas
Gymnopleurus
Gyronotus
Hammondantus
Hamonthophagus
Hansreia
Haroldius
Helictopleurus
Heliocopris
Heteroclitopus
Heteronitis
Holocanthon
Holocephalus
Homalotarsus
Homocopris
Homocorpis
Hyalonthophagus
Hypocanthidium
Ignambia
Isocopris
Ixodina
Janssensantus
Janssensellus
Jossonthophagus
Kheper
Kolbeellus
Krikkenius
Kurtops
Labroma
Larhodius
Latodrepanus
Leotrichillum
Lepanus
Liatongus
Litocopris
Lophodonitis
Macroderes
Malagoniella
Martinezidium
Matthewsius
Megalonitis
Megatharsis
Megathopa
Megathoposoma
Melanocanthon
Mentophilus
Metacatharsius
Microcopris
Milichus
Mimonthophagus
Mnematidium
Mnematium
Monoplistes
Monteithocanthon
Morettius
Namakwanus
Namaphilus
Nanos
Nebulasilvius
Neonitis
Neosaproecius
Neosisyphus
Nesosisyphus
Nesovinsonia
Nitiocellus
Nunoidium
Ochicanthon
Odontoloma
Oficanthon
Oniticellus
Onitis
Onoreidium
Ontherus
Onthobium
Onthophagus
Onychothecus
Oruscatus
Outenikwanus
Oxysternon
Pachylomera
Pachysoma
Panelus
Paracanthon
Parachorius
Paracryptocanthon
Paragymnopleurus
Paraixodina
Paraphacosomoides
Paraphytus
Parascatonomus
Parateuchus
Paroniticellus
Paronthobium
Parvuhowdenius
Peckolus
Pedaria
Pedaridium
Penalus
Pereiraidium
Phaedotrogus
Phalops
Phanaeus
Pinacopodius
Pinacotarsus
Platyonitis
Pleuronitis
Proagoderus
Pseudignambia
Pseudocanthon
Pseudochironitis
Pseudocopris
Pseudonthobium
Pseudopedaria
Pseudosaproecius
Pseuduroxys
Pycnopanelus
Saphobiamorpha
Saphobius
Sarophorus
Sauvagesinella
Scaptocnemis
Scaptodera
Scarabaeolus
Scarabaeus
Scatimus
Scatonomus
Scatrichus
Sceliages
Scybalocanthon
Scybalophagus
Silvaphilus
Silvinha
Sinapisoma
Sinodrepanus
Sisyphus
Stiptocnemis
Stiptopodius
Stiptotarsus
Streblopus
Sukelus
Sulcophanaeus
Sylvicanthon
Synapsis
Tanzanolus
Temnoplectron
Tesserodon
Tesserodoniella
Tetraechma
Theotimius
Thyregis
Tibiodrepanus
Tiniocellus
Tomogonus
Tragiscus
Trichillidium
Trichillum
Tropidonitis
Uenoeus
Unidentis
Uroxys
Versicorpus
Vulcanocanthon
Walterantus
Xenocanthon
Xinidium
Yvescambefortius
Zonocopris
Aclopidae
Aegialiidae
Alloioscarabaeidae
Aphodiidae
Aulonocnemidae
Belohinidae
Bolboceratidae
Cetoniidae
Diphyllostomatidae
Dynastidae
Eremazidae
Euchiridae
Geotrupidae
Glaphyridae
Glaresidae
Hybosoridae
Lucanidae
Melolonthidae
Ochodaeidae
Orphnidae
Pachypodidae
Passalidae
Passalopalpidae
Pleocomidae
Rutelidae
Septiventeridae
Termitotrogidae
Trogidae
EOL Dynamic Hierarchy 1.1
Life
Cellular Organisms
Eukaryota
Opisthokonta
Metazoa
Bilateria
Protostomia
Ecdysozoa
Panarthropoda
Arthropoda
Pancrustacea
Hexapoda
Insecta
Pterygota
Neoptera
Endopterygota
Coleoptera
Polyphaga
Scarabaeiformia
Scarabaeoidea
Scarabaeidae
(this page)
Acanthonitis
Afrodrepanus
Agamopus
Aliuscanthoniola
Allogymnopleurus
Allonitis
Alloscelus
Amietina
Amphiceratodon
Amphistomus
Anisocanthon
Anoctus
Anomiopsoides
Anomiopus
Anonychonitis
Anoplodrepanus
Aphengium
Aphengoecus
Apotolamprus
Aptenocanthon
Apterepilissus
Aptychonitis
Arachnodes
Ateuchetus
Ateuchites
Ateuchus
Attavicinus
Aulacopris
Ausmontins
Avitortor
Baisarabaeus
Baloghonthobium
Bdelyropsis
Bdelyrus
Besourenga
Bohepilissus
Bolbites
Boletoscapter
Bradypodidium
Bubas
Byrrhidium
Caccobiomorphus
Caccobius
Caeconthobium
Cambefortantus
Cambefortius
Canthidium
Canthochilum
Canthodimorpha
Canthon
Canthonella
Canthonidia
Canthonosoma
Canthotrypes
Catharsiocopris
Catharsius
Cephalodesmius
Chalcocopris
Chalconotus
Cheironitis
Chiron
Circellium
Cleptocaccobius
Clypeodrepanus
Copridaspidus
Copris
Coproecus
Coprophanaeus
Coptodactyla
Coptorhina
Cretorabaeus
Cretoscarabaeus
Cryptocanthon
Cyobius
Cyptochirus
Degallieridium
Delopleurus
Deltepilissus
Deltochilum
Deltorhinum
Demarziella
Dendropaemon
Diabroctis
Diastellopalpus
Dichotomius
Dicranocara
Digitonthophagus
Diorygopyx
Disphysema
Dorbignyolus
Drepanocerus
Drepanoplatynus
Drepanopodus
Drogo
Dwesasilvasedis
Endroedyolus
Ennearabdus
Eodrepanus
Epactoides
Epidrepanus
Epilissus
Epirinus
Escarabaeus
Eucranium
Eudinopus
Euoniticellus
Euonthophagus
Eurysternus
Eusaproecius
Eutrichillum
Falsignambia
Feeridium
Frankenbergerius
Garreta
Genieridium
Gilletellus
Glyphoderus
Gromphas
Gymnopleurus
Gyronotus
Hammondantus
Hamonthophagus
Hansreia
Haroldius
Helictopleurus
Heliocopris
Heteroclitopus
Heteronitis
Holcorobeus
Holocanthon
Holocephalus
Homalotarsus
Homocopris
Homocorpis
Hyalonthophagus
Hybosorites
Hypocanthidium
Ignambia
Isocopris
Ixodina
Janssensantus
Janssensellus
Jossonthophagus
Kheper
Kolbeellus
Krikkenius
Kurtops
Labroma
Larhodius
Latodrepanus
Leotrichillum
Lepanus
Liatongus
Lithanomala
Lithoscarabaeus
Litocopris
Lobateuchus
Lophodonitis
Macroderes
Malagoniella
Martinezidium
Matthewsius
Megalonitis
Megatharsis
Megathopa
Megathoposoma
Melanocanthon
Mentophilus
Metacatharsius
Microcopris
Milichus
Mimonthophagus
Mnematidium
Mnematium
Monoplistes
Monteithocanthon
Morettius
Namakwanus
Namaphilus
Nanos
Nebulasilvius
Neonitis
Neosaproecius
Neosisyphus
Nesosisyphus
Nesovinsonia
Nitiocellus
Nunoidium
Ochicanthon
Odontoloma
Oficanthon
Oniticellus
Onitis
Onoreidium
Ontherus
Onthobium
Onthophagus
Onychothecus
Oruscatus
Outenikwanus
Oxysternon
Pachylomera
Pachysoma
Panelus
Paracanthon
Parachorius
Paracryptocanthon
Paragymnopleurus
Paraixodina
Paraphacosomoides
Paraphytus
Parascatonomus
Parateuchus
Paroniticellus
Paronthobium
Parvuhowdenius
Peckolus
Pedaria
Pedaridium
Penalus
Pereiraidium
Phaedotrogus
Phalops
Phanaeus
Pinacopodius
Pinacotarsus
Platyonitis
Pleuronitis
Proagoderus
Proteroscarabaeus
Pseudignambia
Pseudocanthon
Pseudochironitis
Pseudocopris
Pseudonthobium
Pseudopedaria
Pseudosaproecius
Pseuduroxys
Pycnopanelus
Saphobiamorpha
Saphobius
Sarophorus
Sauvagesinella
Scaptocnemis
Scaptodera
Scarabaeolus
Scarabaeus
Scatimus
Scatonomus
Scatrichus
Sceliages
Scybalocanthon
Scybalophagus
Silvaphilus
Silvinha
Sinapisoma
Sinodrepanus
Sisyphus
Stiptocnemis
Stiptopodius
Stiptotarsus
Streblopus
Sukelus
Sulcophanaeus
Sylvicanthon
Synapsis
Tanzanolus
Temnoplectron
Tesserodon
Tesserodoniella
Tetraechma
Theotimius
Thyregis
Tibiodrepanus
Tiniocellus
Tomogonus
Tragiscus
Trichillidium
Trichillum
Tropidonitis
Uenoeus
Unidentis
Uroxys
Versicorpus
Vulcanocanthon
Walterantus
Xenocanthon
Xinidium
Yvescambefortius
Zonocopris
Aclopidae
Aegialiidae
Alloioscarabaeidae
Aphodiidae
Aulonocnemidae
Belohinidae
Bolboceratidae
Cetoniidae
Diphyllostomatidae
Dynastidae
Eremazidae
Euchiridae
Geotrupidae
Glaphyridae
Glaresidae
Hybosoridae
Lucanidae
Melolonthidae
Ochodaeidae
Orphnidae
Pachypodidae
Passalidae
Passalopalpidae
Pleocomidae
Rutelidae
Septiventeridae
Termitotrogidae
Trogidae
EOL Dynamic Hierarchy Version 2.2.4
Scarabaeidae
(this page)
Acanthonitis
Afrodrepanus
Agamopus
Aliuscanthoniola
Allogymnopleurus
Allonitis
Alloscelus
Amietina
Amphiceratodon
Amphistomus
Anisocanthon
Anoctus
Anomiopsoides
Anomiopus
Anonychonitis
Anoplodrepanus
Aphengium
Aphengoecus
Apotolamprus
Aptenocanthon
Apterepilissus
Aptychonitis
Arachnodes
Ateuchetus
Ateuchus
Attavicinus
Aulacopris
Ausmontins
Baloghonthobium
Bdelyropsis
Bdelyrus
Besourenga
Bohepilissus
Bolbites
Boletoscapter
Bradypodidium
Bubas
Byrrhidium
Caccobiomorphus
Caccobius
Caeconthobium
Cambefortantus
Cambefortius
Canthidium
Canthochilum
Canthodimorpha
Canthon
Canthonella
Canthonidia
Canthonosoma
Canthotrypes
Catharsiocopris
Catharsius
Cephalodesmius
Chalcocopris
Chalconotus
Cheironitis
Chiron
Circellium
Cleptocaccobius
Clypeodrepanus
Copridaspidus
Copris
Coproecus
Coprophanaeus
Coptodactyla
Coptorhina
Cryptocanthon
Cyobius
Cyptochirus
Degallieridium
Delopleurus
Deltepilissus
Deltochilum
Deltorhinum
Demarziella
Dendropaemon
Diabroctis
Diastellopalpus
Dichotomius
Dicranocara
Digitonthophagus
Diorygopyx
Disphysema
Dorbignyolus
Drepanocerus
Drepanoplatynus
Drepanopodus
Drogo
Dwesasilvasedis
Endroedyolus
Ennearabdus
Eodrepanus
Epactoides
Epidrepanus
Epilissus
Epirinus
Escarabaeus
Eucranium
Eudinopus
Euoniticellus
Euonthophagus
Eurysternus
Eusaproecius
Eutrichillum
Falsignambia
Feeridium
Frankenbergerius
Garreta
Genieridium
Gilletellus
Glyphoderus
Gromphas
Gymnopleurus
Gyronotus
Hammondantus
Hamonthophagus
Hansreia
Haroldius
Helictopleurus
Heliocopris
Heteroclitopus
Heteronitis
Holocanthon
Holocephalus
Homalotarsus
Homocopris
Homocorpis
Hyalonthophagus
Hypocanthidium
Ignambia
Isocopris
Ixodina
Janssensantus
Janssensellus
Jossonthophagus
Kheper
Kolbeellus
Krikkenius
Kurtops
Labroma
Larhodius
Latodrepanus
Leotrichillum
Lepanus
Liatongus
Litocopris
Lophodonitis
Macroderes
Malagoniella
Martinezidium
Matthewsius
Megalonitis
Megatharsis
Megathopa
Megathoposoma
Melanocanthon
Mentophilus
Metacatharsius
Microcopris
Milichus
Mimonthophagus
Mnematidium
Mnematium
Monoplistes
Monteithocanthon
Morettius
Namakwanus
Namaphilus
Nanos
Nebulasilvius
Neonitis
Neosaproecius
Neosisyphus
Nesosisyphus
Nesovinsonia
Nitiocellus
Nunoidium
Ochicanthon
Odontoloma
Oficanthon
Oniticellus
Onitis
Onoreidium
Ontherus
Onthobium
Onthophagus
Onychothecus
Oruscatus
Outenikwanus
Oxysternon
Pachylomera
Pachysoma
Panelus
Paracanthon
Parachorius
Paracryptocanthon
Paragymnopleurus
Paraixodina
Paraphacosomoides
Paraphytus
Parascatonomus
Parateuchus
Paroniticellus
Paronthobium
Parvuhowdenius
Peckolus
Pedaria
Pedaridium
Penalus
Pereiraidium
Phaedotrogus
Phalops
Phanaeus
Pinacopodius
Pinacotarsus
Platyonitis
Pleuronitis
Proagoderus
Pseudignambia
Pseudocanthon
Pseudochironitis
Pseudocopris
Pseudonthobium
Pseudopedaria
Pseudosaproecius
Pseuduroxys
Pycnopanelus
Saphobiamorpha
Saphobius
Sarophorus
Sauvagesinella
Scaptocnemis
Scaptodera
Scarabaeolus
Scarabaeus
Scatimus
Scatonomus
Scatrichus
Sceliages
Scybalocanthon
Scybalophagus
Silvaphilus
Silvinha
Sinapisoma
Sinodrepanus
Sisyphus
Stiptocnemis
Stiptopodius
Stiptotarsus
Streblopus
Sukelus
Sulcophanaeus
Sylvicanthon
Synapsis
Tanzanolus
Temnoplectron
Tesserodon
Tesserodoniella
Tetraechma
Theotimius
Thyregis
Tibiodrepanus
Tiniocellus
Tomogonus
Tragiscus
Trichillidium
Trichillum
Tropidonitis
Uenoeus
Unidentis
Uroxys
Versicorpus
Vulcanocanthon
Walterantus
Xenocanthon
Xinidium
Yvescambefortius
Zonocopris
Aclopidae
Aegialiidae
Alloioscarabaeidae
Aphodiidae
Aulonocnemidae
Belohinidae
Bolboceratidae
Cetoniidae
Diphyllostomatidae
Dynastidae
Eremazidae
Euchiridae
Geotrupidae
Glaphyridae
Glaresidae
Hybosoridae
Lucanidae
Melolonthidae
Ochodaeidae
Orphnidae
Pachypodidae
Passalidae
Passalopalpidae
Pleocomidae
Rutelidae
Septiventeridae
Termitotrogidae
Trogidae
GBIF classification
Animalia
Arthropoda
Insecta
Coleoptera
Euchiridae
(this page)
Cheirotonus Hope 1840
Euchirus Kirby & Spence 1828
Propomacrus Newman 1837
Aderidae
Agapythidae
Agyrtidae
Alexiidae
Amphizoidae
Anomiopsis Westwood 1837
Anthribidae
Aphodiidae
Apionidae
Apterogena Ardoin 1962
Archeocrypticidae
Artematopidae
Artematopodidae
Attelabidae
Aulonocnemidae
Belidae
Boganiidae
Boridae
Brentidae
Bruchidae
Buprestidae
Byturidae
Callirhipidae
Carabidae
Cavognathidae
Cebrionidae
Cephaloidae
Ceratocanthidae
Cerophytidae
Cetoniidae
Chaetosomatidae
Chalcodryidae
Ciidae
Cleridae
Cneoglossidae
Colydiidae
Coptoclavidae
Corylophidae
Curculionidae
Dasyceridae
Decliniidae
Diphyllostomatidae
Discolomidae
Drilidae
Dynastidae
Fatua Crotch 1876
Gerardia Pic 1954
Glaphyridae
Glaresidae
Glauconia Paulian 1939
Harpalidium Westwood 1854
Helotidae
Hobartiidae
Hologymnia Schoch 1895
Homalisidae
Hybosoridae
Hygrobiidae
Lamingtoniidae
Lathridiidae
Lepiceridae
Leptinidae
Lucanidae
Malachiidae
Monommidae
Murmidiidae
Myraboliidae
Omalisidae
Omocera Blanchard 1846
Oxydactylus Martins 1965
Paragyrinus Handlirsch 1906
Passalidae
Perimylopidae
Phengodidae
Phloeostichidae
Plastoceridae
Platypodidae
Plavichydissus Pic 1946
Priasilphidae
Protocucujidae
Pseudoheterochelus wallekraalensis
Pterogeniidae
Raymondionymidae
Rhadalidae
Rhizophagidae
Ripiphoridae
Scalopoides Bode 1953
Schizopodidae
Scolytidae
Sparmannia Laporte 1840
Sphaerosomatidae
Synteliidae
Syntomopterus Bode 1953
Tasmosalpingidae
Termitotrogidae
Thurmannia Heer 1852
Thymalidae
Torridincolidae
Trachelostenidae
Trictenotomidae
Ulodidae
579 additional siblings truncated for brevity.
See the
resource file
for a full list.
GBIF classification
Animalia
Arthropoda
Insecta
Coleoptera
Scarabaeidae
(this page)
Acanthonitis Janssens 1937
Afrodrepanus Krikken 2009
Agamopus Bates 1887
Aliuscanthoniola Deschodt & Scholtz 2008
Allogymnopleurus Janssens 1940
Allonitis Janssens 1936
Alloscelus Boucomont 1923
Amietina Cambefort 1981
Amphiceratodon Huchet 2000
Amphistomus van Lansberge 1874
Anisocanthon Martínez & Pereira 1956
Anoctus Sharp 1875
Anomalacra Casey 1915
Anomiopsoides Blackwelder 1944
Anomiopus Westwood 1842
Anonychonitis Janssens 1950
Anoplodrepanus
Anoplosiagum Blanchard 1850
Aphengium Harold 1868
Aphengoecus Péringuey 1901
Aphodius
Apotolamprus Olsoufieff 1947
Aptenocanthon Matthews 1974
Apterepilissus Montreuil 2011
Aptychonitis Janssens 1937
Arachnodes Westwood 1843
Ateuchetus Bedel 1892
Ateuchites Meunier 1898
Ateuchus Weber 1801
Attavicinus Philips & Bell 2008
Aulacopris White 1859
Ausmontins Deschodt & Davis 2018
Avitortor Ponomarenko 1977
Baisarabaeus Nikolajev 2005
Baloghonthobium Paulian 1986
Bdelyropsis Pereira, Vulcano & Martínez 1960
Bdelyrus Harold 1869
Besourenga
Bohepilissus Paulian 1975
Bolbites Harold 1868
Boletoscapter Matthews 1974
Bolobochromus meridionalis Krikken 1977
Bradypodidium
Bubas Mulsant 1842
Byrrhidium Harold 1869
Caccobiomorphus Balthasar 1964
Caccobius Thomson 1859
Caeconthobium Paulian 1984
Calhyboma Kolbe 1893
Callistemous intrusus
Cambefortantus Paulian 1986
Cambefortius Branco 1989
Canthidium Erichson 1847
Canthochilum Chapin 1934
Canthodimorpha Davis, Scholtz & Harrison 1999
Canthon Hoffmannsegg 1817
Canthonella Chapin 1930
Canthonidia Paulian 1938
Canthonosoma Mac Leay 1871
Canthotrypes Paulian 1939
Catharsiocopris Balthasar 1967
Catharsius Hope 1837
Cephalodesmius Westwood 1842
Cetonischema Reitter 1899
Chalcocopris Burmeister 1846
Chalconotus Dejean 1833
Cheironitis Lansberge 1875
Chiron Macleay 1819
Chrysopotosia Miksic 1966
Circellium Latreille 1829
Cleptocaccobius Cambefort 1984
Clypeodrepanus Krikken 2009
Coelodera
Copridaspidus Boucomont 1920
Copris Geoffroy 1762
Coproecus Reiche 1841
Coprophanaeus D' Olsoufieff 1924
Coptodactyla Burmeister 1846
Coptorhina Hope 1833
Costleya Broun 1893
Cratocnemus Blanchard 1842
Cretaegialia Nikolayev 1994
Cretomelolontha Nikolajev 1998
Cretorabaeus Nikolajev 1995
Cretoscarabaeus Nikolajev 1995
Cretoserica Nikolajev 1998
Cryptocanthon Balthasar 1942
Ctilocephala Burmeister 1855
Cyobius Sharp 1875
Cyptochirus Lesne 1900
Degallieridium
Delopleurus Erichson 1847
Deltepilissus Silvério Pereira 1949
Deltochilum Eschscholtz 1822
Deltohyboma Martinez 1947
Deltorhinum Harold 1869
Demarziella Balthasar 1961
Dendropaemon Perty 1830
Diabroctis Gistel 1857
Diastellopalpus Lansberge 1886
Diathermus Kraatz 1897
Dichotomius Hope 1838
Dicranobia Reitter 1900
Dicranocara Frolov & Scholtz 2003
Dicronorhina Hope 1837
Digitonthophagus Balthasar 1959
Diorygopyx Matthews 1974
Disphysema Harold 1873
Dorbignyolus Branco 1989
Drepanacanthoides Schmidt 1913
Drepanocerus Kirby 1828
Drepanoplatynus Boucomont 1921
Drepanopodus Janssens 1940
Drogo
Dwesasilvasedis Deschodt & Scholtz 2008
Endroedyolus Scholtz & Howden 1987
Ennearabdus Van Lansberge 1874
Eodrepanus Barbero, Palestrini & Roggero 2009
Epactoides Olsoufieff 1947
Epidrepanus Roggero, Barbero & Palestrini 2015
Epilissus Reiche 1841
Epirinus Reiche 1841
Escarabaeus Zídek & Pokorný 2011
Eucetonia Schoch 1894
Eucranium Brullé 1834
Eudinopus Burmeister 1840
Euglypta Mohnike 1873
Euoniticellus Janssens 1953
Euonthophagus Balthasar 1959
Eusaproecius Branco 1989
Eutrichillum Martinez 1968
Exantichira Soula 1998
Falsignambia Paulian 1987
Feeridium Vaz de Mello 2008
Frankenbergerius Balthasar 1938
Garreta Janssens 1940
Genieridium
Gilletellus Janssens 1937
Glaphyrocanthon Martínez 1948
Glyphoderus Westwood 1837
Gromphas Brullé 1839
Gymnopleurus Illiger 1803
Gyronotus Cameron 1948
Haematonotus Kraatz 1880
Hammondantus Cambefort 1978
Hamonthophagus Roggero, Dierkens, Barbero & Palestrini 2017
Hansreia Halffter & Martinez 1977
Haroldius Boucomont 1914
Helictopleurus Orbigny 1915
Heliocopris Hope 1837
Heteroclitopus Péringuey 1901
Heteronitis Gillet 1911
Heteropelidnota Ohaus 1912
Heterorrhina
Holcorobeus Nikritin 1977
Holocanthon Martínez & Pereira 1956
Holocephalus Hope 1838
Homalotarsus Janssens 1932
Homocopris Burmeister 1846
Homocorpis
Hyalonthophagus Palestrini & Giacone 1988
Hybosorites Nikolajev 1996
Hypocanthidium Balthasar 1938
Ignambia Heller 1916
Ischnocelis
Isocopris Pereira & Martinez 1960
Ixodina Roth 1851
Janssensantus
Janssensellus Cambefort 1976
Jossonthophagus Dierkens, Moretto & Génier 2017
Kheper Janssens 1940
Kolbeellus Jacobson 1906
Krikkenius Branco 1992
Kurtops Roggero, Barbero & Palestrini 2016
Labroma Sharp 1873
Lachnosterna Hope 1837
Larhodius
Latodrepanus Krikken 2009
Leotrichillum
Lepanus Balthasar 1966
Lepesmonyx Paulian 1991
Liatongus Reitter 1893
Lithanomala Nikolajev 1992
Lithoscarabaeus Nikolajev 1992
Litocopris Waterhouse 1891
Litolontha simplex
Lobateuchus Montreuil, Génier & Nel 2010
Lobidion Genier 2010
Lophodonitis Janssens 1938
Lophorrhinides
Macroderes Westwood 1842
Macrophylla Hope 1837
Malagoniella Martínez 1961
Martinezidium
Matthewsius
Mecaspidius Bourgoin 1921
Mecinonota Kraatz 1892
Megalonitis Janssens 1937
Megalorhina Westwood 1847
Megasybacodes Kakizoe, Maruyama & Masumoto 2019
Megatharsis Waterhouse 1891
Megathopa Eschscholtz 1822
Megistophyllas
Melanocanthon Halffter 1958
Mentophilus Laporte de Castelnau 1840
Metacatharsius Paulian 1939
Micrelaphinis Schoch 1896
Microcamenta Brenske 1898
Microcopris Balthasar 1958
Microthyrea Kraatz 1882
Milichus
Miltotrogus Reitter 1902
Mimeoma Casey 1915
Mimonthophagus Balthasar 1963
Mnematidium Ritsema 1888
Mnematium Macleay 1821
Monoplistes Lansberge 1874
Monteithocanthon
Morettius Roggero, Dierkens, Barbero & Palestrini 2017
Namakwanus Scholtz & Howden 1987
Namaphilus
Nanos Westwood 1842
Nebulasilvius Deschodt & Scholtz 2008
Neocanthidium
Neonitis Péringuey 1901
Neosaproecius Branco 1990
Neosisyphus Müller 1942
Neptunides Thomson 1879
Nesosisyphus Vinson 1946
Nesovinsonia Martínez & Pereira 1959
Nitiocellus Branco 2010
Nunoidium
Ochicanthon
Odontoloma Boheman 1857
Oficanthon Paulian 1985
Oncosterna Thomson 1880
Oniticellus Dejean 1821
Onitis Fabricius 1798
Onoreidium
Ontherus Erichson 1847
Onthobium Reiche 1860
Onthocharis Westwood 1847
Onthophagus Latreille 1802
Onychosophrops
Onychothecus Boucomont 1912
Oruscatus Bates 1870
Outenikwanus Scholtz & Howden 1987
Oxycetonia Arrow 1910
Oxygrylius Casey 1915
Oxysternon Laporte de Castelnau 1840
Pachylomera Kirby 1832
Pachysoma Macleay 1821
Panelus Lewis 1895
Paracanthon Balthasar 1938
Parachorius Harold 1873
Paracryptocanthon Howden & Cook 2002
Paragymnopleurus Shipp 1897
Paraixodina
Paraphacosomoides
Paraphytus Harold 1877
Parascatonomus Paulian 1932
Parateuchus Shipp 1895
Parisoleoides Morón 1988
Paroniticellus Balthasar 1963
Paronthobium Paulian 1984
Parvuhowdenius Deschodt & Scholtz 2008
Peckolus Scholtz & Howden 1987
Pedaria Laporte 1832
Pedaridium Harold 1868
Penalus Paulian 1985
Pereiraidium
Phalops Erichson 1847
Phanaeus Mac Leay 1819
Phycocus Broun 1883
Piastellopalpus guingnedens
Pinacopodius Branco 1992
Pinacotarsus Harold 1875
Plagiochilus Wasmann 1900
Platyonitis Janssens 1942
Pleuronitis Van Lansberge 1875
Plusiotus
Poecilodiscus Broun 1895
Prigenia Mohnike 1871
Proagoderus Lansberge 1883
Proteroscarabaeus Grabau 1923
Protopalnia Nikolajev 2007
Pseudeuryomia Kraatz 1894
Pseudignambia Paulian & Pluot-Sigwalt 1985
Pseudoapterogyna Martínez De La Escalera 1914
Pseudocanthon Bates 1887
Pseudocopris Ferreira 1960
Pseudomecinonota Mikšic 1972
Pseudonthobium Paulian 1984
Pseudopedaria Felsche 1904
Pseudosaproecius Balthasar 1941
Pseuduroxys Balthasar 1938
Pycnopanelus Arrow 1931
Rhinacosmus Kraatz 1895
Rhomborrhina
Rimulopia alluaudi Lacroix 1997
Ruterielina Rojkoff 2010
Sabatinellopia ovata Lacroix 1998
Saphobiamorpha Brookes 1944
Saphobius Sharp 1873
Sarophorus Erichson 1847
Sauvagesinella Paulian 1935
Scaptocnemis Péringuey 1901
Scaptodera Hope 1837
Scarabaeolus Balthasar 1965
Scarabaeus
Scatimus Erichson 1847
Scatonomus Erichson 1835
Scatrichus Genier & Kohlmann 2003
Sceliages Westwood 1837
Scybalocanthon Martínez 1948
Scybalophagus
Silvaphilus Roets & Oberlander 2010
Silvinha
Sinapisoma Boucomont 1928
Sinodrepanus Simonis 1985
Sisyphus Latreille 1807
Stenovalgus Kolbe 1892
Stiptocnemis Branco 1989
Stiptopodius Harold 1871
Stiptotarsus Branco 1989
Streblopus Lansberge 1874
Sukelus Branco 1992
Sulcophanaeus Olsoufieff 1924
Sylvicanthon Halffter & Martínez 1977
Synapsis Bates 1868
Taiwanoserica
Temnoplectron Westwood 1842
Temnorrhynchus Hope 1837
Tesserodon Hope 1837
Tesserodoniella
Tetraechma Blanchard 1841
Thaumastopeus Kraatz 1885
Theotimius Huchet 2000
Thyregis Blackburn 1904
Tibiodrepanus Krikken 2009
Tiniocellus Péringuey 1902
Tomogonus D' Orbigny 1904
Tragiscus Klug 1855
Trichelasmus Sharp 1877
Trichillidium
Trichillum Harold 1868
Trichiomorphus Bourgoin 1919
Trinodia Casey 1915
Uenoeus
Unidentis Josso & Prévost 2008
Uroxys Westwood 1842
Versicorpus Deschodt, Davis & Scholtz 2011
Vulcanocanthon Pereira & Martinez 1960
Walterantus Cambefort 1977
Xalpirta Skelley & Cekalovic 2001
Xenocanthon Martínez 1952
Xinidium Harold 1869
Yvescambefortius Ochi & Kon 1996
Zonocopris Arrow 1932
Aderidae
Agapythidae
Agyrtidae
Alexiidae
Amphizoidae
Anomiopsis Westwood 1837
Anthribidae
Aphodiidae
Apionidae
Apterogena Ardoin 1962
Archeocrypticidae
Artematopidae
Artematopodidae
Attelabidae
Aulonocnemidae
Belidae
Boganiidae
Boridae
Brentidae
Bruchidae
Buprestidae
Byturidae
Callirhipidae
Carabidae
Cavognathidae
Cebrionidae
Cephaloidae
Ceratocanthidae
Cerophytidae
Cetoniidae
Chaetosomatidae
Chalcodryidae
Ciidae
Cleridae
Cneoglossidae
Colydiidae
Coptoclavidae
Corylophidae
Curculionidae
Dasyceridae
Decliniidae
Diphyllostomatidae
Discolomidae
Drilidae
Dynastidae
Euchiridae
Fatua Crotch 1876
Gerardia Pic 1954
Glaphyridae
Glaresidae
Glauconia Paulian 1939
Harpalidium Westwood 1854
Helotidae
Hobartiidae
Hologymnia Schoch 1895
Homalisidae
Hybosoridae
Hygrobiidae
Lamingtoniidae
Lathridiidae
Lepiceridae
Leptinidae
Lucanidae
Malachiidae
Monommidae
Murmidiidae
Myraboliidae
Omalisidae
Omocera Blanchard 1846
Oxydactylus Martins 1965
Paragyrinus Handlirsch 1906
Passalidae
Perimylopidae
Phengodidae
Phloeostichidae
Plastoceridae
Platypodidae
Plavichydissus Pic 1946
Priasilphidae
Protocucujidae
Pseudoheterochelus wallekraalensis
Pterogeniidae
Raymondionymidae
Rhadalidae
Rhizophagidae
Ripiphoridae
Scalopoides Bode 1953
Schizopodidae
Scolytidae
Sparmannia Laporte 1840
Sphaerosomatidae
Synteliidae
Syntomopterus Bode 1953
Tasmosalpingidae
Termitotrogidae
Thurmannia Heer 1852
Thymalidae
Torridincolidae
Trachelostenidae
Ulodidae
579 additional siblings truncated for brevity.
See the
resource file
for a full list.
Integrated Taxonomic Information System (ITIS) Taxonomic Hierarchy
Animalia
Bilateria
Protostomia
Ecdysozoa
Arthropoda
Hexapoda
Insecta
Pterygota
Neoptera
Holometabola
Coleoptera Linnaeus 1758
Polyphaga Emery 1886
Scarabeiformia Crowson 1960
Scarabaeoidea Latreille 1802
Scarabaeidae Latreille 1802
(this page)
Aclopinae Blanchard 1850
Allidiostomatinae Arrow 1940
Aphodiinae Leach 1815
Cetoniinae Leach 1815
Dynamopodinae Arrow 1911
Dynastinae MacLeay 1819
Euchirinae Hope 1840
Melolonthinae Leach 1819
Orphninae Erichson 1847
Pachypodinae Erichson 1840
Phaenomeridinae Erichson 1847
Rutelinae MacLeay 1819
Scarabaeinae Latreille 1802
Belohinidae Paulian 1959
Diphyllostomatidae Holloway 1972
Geotrupidae Latreille 1802
Glaphyridae MacLeay 1819
Glaresidae Kolbe 1905
Hybosoridae Erichson 1847
Lucanidae Latreille 1804
Ochodaeidae Mulsant & Rey 1871
Passalidae Leach 1815
Pleocomidae Le Conte 1861
Trogidae MacLeay 1819
IUCN
Animalia
Arthropoda
Insecta
Coleoptera
Scarabaeidae
(this page)
Aegialia concinna Gordon & Cartwright 1977
Aegialia crescenta Gordon & Cartwright 1977
Afrodrepanus marshalli (Boucomont 1921)
Allogymnopleurus chloris Klug 1855
Allogymnopleurus histrio Balthasar 1963
Allogymnopleurus thalassinus Klug 1855
Amietina larrochei Cambefort
Amphistomus complanatus Matthews 1974
Amphistomus speculifer Matthews 1974
Anisocanthon pygmaeus (Gillet 1911)
Anomiopus ataenioides (Martinez 1952)
Anomiopus birai Canhedo 2006
Anomiopus bonariensis (Bruch 1925)
Anomiopus caputipilus Canhedo 2004
Anomiopus gracilis Canhedo 2006
Anomiopus howdeni Canhedo 2006
Anomiopus idei Canhedo 2006
Anomiopus lacordairei (Waterhouse 1891)
Anomiopus laetus (Waterhouse 1891)
Anomiopus nigrocoeruleus (Martinez 1955)
Anomiopus parallelus (Harold 1862)
Anomiopus pictus (Harold 1862)
Anomiopus smaragdinus (Westwood 1842)
Anomiopus soledari Canhedo 2004
Anomiopus virescens (Westwood 1842)
Anonychonitis freyi Janssens
Aptenocanthon hopsoni (Carter 1936)
Aptenocanthon winyar Storey & Monteith 2000
Ateuchetus cicatricosus (Lucas 1846)
Ateuchetus laticollis Linnaeus 1767
Ateuchetus puncticollis Latreille 1819
Ateuchetus semipunctatus (Fabricius 1792)
Ateuchetus variolosus (Fabricius 1787)
Ateuchus ambiguus Martinez & Martinez 1990
Ateuchus carbonarius (Harold 1868)
Ateuchus contractus (Balthasar 1939)
Ateuchus ecuadorensis (Boucomont 1928)
Ateuchus euchalceus (Balthasar 1939)
Ateuchus laevicollis (Harold 1868)
Ateuchus latus (Boucomont 1928)
Ateuchus murrayi (Harold 1868)
Ateuchus puncticolle (Harold 1867)
Ateuchus romani (Boucomont 1927)
Ateuchus squalidus (Fabricius 1775)
Aulacopris matthewsi Storey 1986
Aulacopris reichei White 1859
Bdelyrus apaporisae Cook 1998
Bdelyrus boliviensis Cook 1998
Bdelyrus cochabambae Cook 2000
Bdelyrus grandis Cook 1998
Bdelyrus howdeni Cook 1998
Bdelyrus iquitosensis Cook 2000
Bdelyrus leptomerus Cook 1998
Bdelyrus metaensis Cook 1998
Bdelyrus paraensis Cook 1998
Bdelyrus triangulus Cook 1998
Besourenga horacioi (Martínez 1967)
Boletoscapter furcatus Matthews 1974
Bubas bison (Linnaeus 1767)
Bubas bubaloides Janssens 1938
Bubas bubalus (Olivier 1811)
Byrrhidium convexum Scholtz & Howden 1987
Caccobius cavatus D' Orbigny 1908
Caccobius croceocinctus D' Orbigny 1913
Caccobius ferrugineus (Fåhraeus 1857)
Caccobius nigritulus Klug 1855
Canthidium abbreviatum Harold 1867
Canthidium angulicolle Balthasar 1939
Canthidium aterrimum Harold 1867
Canthidium atomarium Balthasar 1939
Canthidium aurichalceum Preudhomme de Borre 1886
Canthidium bicolor Boucomont 1928
Canthidium bokermanni (Martinez, Halffter & Pereira 1964)
Canthidium calidum Harold 1880
Canthidium cavifrons Balthasar 1939
Canthidium coerulescens Balthasar 1939
Canthidium deplanatum Balthasar 1939
Canthidium erythropterum (Lucas 1857)
Canthidium femoratum Boucomont 1935
Canthidium flabellatum Harold 1883
Canthidium flavicorne (Blanchard 1846)
Canthidium flavipes Harold 1867
Canthidium flavum Balthasar 1939
Canthidium glabricolle Harold 1867
Canthidium guyanense Boucomont 1928
Canthidium haagi Harold 1867
Canthidium latum (Blanchard 1846)
Canthidium luteum Balthasar 1939
Canthidium magnum Harold 1871
Canthidium manni Arrow 1913
Canthidium metallicum Harold 1867
Canthidium nitidum (Blanchard 1846)
Canthidium opacum Balthasar 1939
Canthidium sladeni Arrow 1903
Canthidium subdopuncticolle Howden & Young 1981
Canthidium viride (Lucas 1857)
Canthidium viridicolle (Blanchard 1846)
Canthidium viridiobscurum Boucomont 1928
Canthochilum anacaona Zayas & Matthews 1966
Canthon angularis Harold 1868
Canthon brunnipennis Schmidt 1922
Canthon coloratus Schmidt 1922
Canthon columbianus Schmidt 1920
Canthon corpulentus Harold 1868
Canthon corruscans Castelnau 1840
Canthon daguerrei Martinez 1951
Canthon deplanatus Harold 1868
Canthon divinator Gistel 1857
Canthon edentulus Harold 1868
Canthon formosus Harold 1868
Canthon gemellatus Erichson 1847
Canthon helleri Schmidt 1922
Canthon heyrovskyi Balthasar 1939
Canthon janthinus Blanchard 1843
Canthon laminatus Balthasar 1939
Canthon lituratus (Germar 1813)
Canthon lividus Blanchard 1846
Canthon luctuosus Harold 1868
Canthon lunatus Schmidt 1922
Canthon mutabilis Lucas 1857
Canthon orfilai Martinez 1949
Canthon ornatus Redtenbacher 1867
Canthon plagiatus Harold 1880
Canthon podagricus Harold 1868
Canthon quadripunctatus Redtenbacher 1867
Canthon quinquemaculatus Castelnau 1840
Canthon rubrescens Blanchard 1846
Canthon rutilans Harold 1868
Canthon septemmaculatus Latreille 1811
Canthon sericatus Schmidt 1922
Canthon simulans (Martinez 1950)
Canthon smaragdulus (Fabricius 1781)
Canthon sordidus Harold 1868
Canthon subhyalinus Harold 1867
Canthon tetraodon Blanchard 1846
Canthon triangularis (Drury 1770)
Canthon unicolor Balthasar 1939
Canthon virens Mannerheim 1829
Canthon vulcanoae Pereira & Martinez 1956
Canthonella gomezi (Halffter & Martinez 1968)
Canthonella instriata (Boucomont 1928)
Catharsius bradshawi Lansberge 1887
Catharsius brittoni Ferreira 1963
Catharsius calaharicus Kolbe 1893
Catharsius chinai Ferreira 1960
Catharsius oedipus Felsche 1907
Catharsius pandion Harold 1877
Catharsius rhinoceros Klug 1855
Cephalodesmius armiger Westwood 1842
Cephalodesmius laticollis Pascoe 1863
Cheironitis audens (Péringuey 1901)
Cheironitis furcifer (Rossi 1792)
Cheironitis indicus Lansberge 1875
Cheironitis scabrosus (Fabricius 1776)
Cheirotonus jambar Kurosawa 1984
Clypeodrepanus striatus (Boucomont 1921)
Copris amabilis Kolbe 1914
Copris armiger Gillet 1910
Copris capensis Waterhouse 1891
Copris complexus Nguyen-phung 1989
Copris fallaciosus Gillet 1907
Copris fallax Felsche 1910
Copris fidius (Olivier 1789)
Copris inhalatus Quedenfeldt 1884
Copris morphaeus Gillet 1932
Copris pueli Mollandin de Boissy 1905
Copris ritsemae Harold 1875
Copris serius Nguyen-phung 1987
Copris umbilicatus Abeille de Perrin 1901
Copris victorini Boheman 1857
Copris wiesei Kolbe 1914
Coproecus hemisphaericus Guerin-Meneville 1844
Coprophanaeus corythus (Harold 1863)
Coprophanaeus gamezi Arnaud 2002
Coprophanaeus ohausi (Felsche 1911)
Coprophanaeus pertyi (d'Olsufieff 1924)
Coprophanaeus rigoutorum Arnaud 2002
Coprophanaeus terrali Arnaud 2002
Coptodactyla lesnei Paulian 1933
Coptodactyla nitida Paulian 1933
Coptodactyla storeyi Reid 2000
Coptorhina nitefacta Gillet 1932
Cryptocanthon altus Howden 1976
Cryptocanthon andersoni Cook 2002
Cryptocanthon campbellorum Howden 1973
Cryptocanthon foveatus Cook 2002
Cryptocanthon genieri Cook 2002
Cryptocanthon medinae Cook 2002
Cryptocanthon nebulinus Howden 1973
Cryptocanthon otonga Cook 2002
Cryptocanthon punctatus Cook 2002
Cyptochirus ambiguus (Kirby 1828)
Deltochilum batesi Paulian 1938
Deltochilum brasiliense Castelnau 1840
Deltochilum carinatum (Westwood 1837)
Deltochilum cristinae (Martínez 1991)
Deltochilum guyanense Paulian 1933
Deltochilum icarus (Olivier 1789)
Deltochilum peruanum Paulian 1938
Deltochilum rubripenne Gory 1831
Deltochilum violetae (Martínez 1991)
Deltorrhinum batesi Harold 1867
Demarziella eungella Matthews 1976
Demarziella imitatrix Balthasar 1967
Demarziella planitarsis Petrovitz 1971
Demarziella storeyi Matthews 1976
Demarziella tropicalis Matthews 1976
Dendropaemon crenatostriatus Felsche 1909
Dendropaemon tenuitarsis Felsche 1909
Diastellopalpus infernalis (Lansberge 1887)
Diastellopalpus thomsoni (Bates 1888)
Dichotomius alyattes (Harold 1880)
Dichotomius apicalis (Luederwaldt 1931)
Dichotomius calcaratus Arrow 1913
Dichotomius crinicollis (Germar 1824)
Dichotomius dahli (Landin 1956)
Dichotomius eucranioides Pereira & Andretta 1955
Dichotomius fallax (Harold 1880)
Dichotomius glaucus (Harold 1869)
Dichotomius horridus (Felsche 1911)
Dichotomius nimuendaju Luederwaldt 1925
Dichotomius nutans (Harold 1867)
Dichotomius planicollis (Gillet 1911)
Dichotomius podalirius (Felsche 1901)
Dichotomius prietoi Martinez & Martinez 1982
Dichotomius problematicus (Luederwaldt 1924)
Dichotomius punctulatipennis (Luederwaldt 1930)
Dichotomius schiffleri
Dichotomius tristis (Luederwaldt 1924)
Dichotomius worontzowi (Pereira 1942)
Dicranocara deschodti Frolov & Scholtz 2003
Diorygopyx duplodentatus Matthews 1974
Diorygopyx niger Matthews 1974
Endroedyolus paradoxus Scholtz & Howden 1987
Eodrepanus fastiditus (Péringuey 1901)
Eodrepanus parallelus (Raffray 1877)
Epirinus asper Péringuey 1901
Epirinus comosus Péringuey 1901
Epirinus gratus Péringuey 1901
Epirinus relictus Scholtz & Howden 1987
Epirinus sulcipennis Boheman 1857
Epirinus validus Péringuey 1901
Euoniticellus inaequalis (Reiche 1850)
Euoniticellus kawanus Janssens 1939
Euoniticellus nasicornis (Reiche 1849)
Euoniticellus perniger Cambefort & Mauchamps 1980
Euonthophagus crocatus (Mulsant & Godart 1870)
Euonthophagus maindroni (D' Orbigny 1897)
Euonthophagus tissoni (Reitter 1906)
Frankenbergerius armatus (Boheman 1857)
Frankenbergerius forcipatus (Harold 1880)
Frankenbergerius nitidus Frolov & Scholtz 2005
Frankenbergerius opacus Frolov & Scholtz 2005
Garreta azureus (Fabricius 1801)
Garreta malleolus (Kolbe 1895)
Garreta nitens (Olivier 1789)
Genieridium medinae
Gromphas aeruginosa (Perty 1830)
Gromphas amazonica Bates 1870
Gymnopleurus aenescens Wiedemann 1821
Gymnopleurus aeruginosus Harold 1867
Gymnopleurus andreaei Ferreira 1954
Gymnopleurus bicolor Latreille 1823
Gymnopleurus humanus Macleay 1821
Gymnopleurus reichei Waterhouse 1890
Gymnopleurus sturmi MacLeay 1821
Gymnopleurus thelwalli Waterhouse 1890
Gymnopleurus virens Erichson 1843
Gyronotus dispar (Felsche 1911)
Gyronotus mulanjensis Davis, Scholtz & Harrison 1999
Gyronotus pumilus (Boheman 1857)
Hansreia affinis (Fabricius 1801)
Heliocopris faunus Boheman 1857
Heliocopris japetus Klug 1855
Heteronitis castelnaui (Harold 1862)
Holocephalus sculptus (Gillet 1907)
Homocopris torulosus (Eschscholtz 1822)
Isocopris tarsalis Luederwaldt 1931
Ixodina freyi (Janssens 1953)
Kolbeellus ateuchoides Lansberge 1875
Latodrepanus laticollis (Fahraeus 1857)
Lepanus bidentatus Wilson 1922
Lepanus gelasinus Matthews 1974
Lepanus pisoniae Lea 1923
Lepanus politus Carter 1936
Lepanus pygmaeus Macleay 1888
Lepanus ustulatus Lansberge 1874
Lepanus villosus Matthews 1974
Liatongus arrowi Boucomont 1921
Liatongus urus Kolbe 1914
Macroderes cornutus Frolov & Scholtz 2004
Macroderes endroedyi Frolov & Scholtz 2004
Macroderes undulatus Preudhomme de Borre 1880
Malagoniella argentina (Gillet 1911)
Malagoniella chalybaea (Blanchard 1846)
Matthewsius illawarrensis (Matthews 1974)
Matthewsius rossi (Matthews 1974)
Megalonitis bohemani (van Lansberge 1875)
Mellissius adumbratus Wollaston 1869
Mellissius eudoxus Wollaston 1877
Mellissius oryctoides Decelle 1972
Metacatharsius inermis Castelnau 1840
Metacatharsius pumilioniformis Ferreira 1964
Metacatharsius transvaalensis Balthasar 1968
Mimonthophagus limbibasis (d'Orbigny 1905)
Monoplistes occidentalis (Macleay 1888)
Monoplistes phanophilus Lea 1923
Monoplistes tropicus Lea 1923
Namakwanus streyi Frolov 2005
Neosisyphus fortuitus Peringuey 1901
Neosisyphus kuhni Haaf 1955
Neosisyphus macrorubrus (Paschalidis 1974)
Neosisyphus mirabilis Arrow 1927
Neosisyphus rubrus (Paschalidis 1974)
Odontoloma endroedyi Howden & Scholtz 1987
Odontoloma louwi Howden & Scholtz 1987
Odontoloma obscurum Howden & Scholtz 1987
Odontoloma pygidiale Peringuey 1901
Oniticellus pseudoplanatus Balthasar 1964
Onitis adelphus Kolbe 1914
Onitis aerarius Harold 1878
Onitis aeruginosus Klug 1855
Onitis belial Fabricius 1798
Onitis caffer Boheman 1857
Onitis dispar Péringuey 1896
Onitis ezechias Reiche 1856
Onitis fulgidus Klug 1855
Onitis granulisetosus Ferreira 1976
Onitis ion (Olivier 1789)
Onitis keniensis Gillet 1933
Onitis kingstoni Krikken 1977
Onitis laminosus Ferreira 1976
Onitis mendax Gillet 1918
Onitis meyeri Kolbe 1891
Onitis mniszechi Lansberge 1875
Onitis monstrosus Felsche 1910
Onitis nemoralis Gillet 1932
Onitis obscurus Lansberge 1886
Onitis parainflaticollis Ferreira 1977
Onitis parvulus Balthasar 1963
Onitis retrodentatus Gillet 1932
Onitis robustus Boheman 1857
Onitis tanzaniensis Ferreira 1977
Onitis uncinatus Klug 1855
Onoreidium howdeni (Ferreira & Galileo 1993)
Ontherus aequatorius Bates 1891
Ontherus androgynus Génier 1996
Ontherus azteca Harold 1869
Ontherus brevicollis Kirsch 1870
Ontherus dentatus Luederwaldt 1930
Ontherus erosioides Luederwaldt 1930
Ontherus grandis Luederwaldt 1931
Ontherus hadros Génier 1996
Ontherus irinus Balthasar 1938
Ontherus laminifer Balthasar 1938
Ontherus obliquus Génier 1996
Ontherus raptor Génier 1996
Ontherus virescens (Lucas 1859)
Onthophagus adelaidae Hope 1846
Onthophagus aequepubens D' Orbigny 1905
Onthophagus aethiopicus D' Orbigny 1902
Onthophagus albarracinus Baraud 1979
Onthophagus alquirta Matthews 1972
Onthophagus amphioxus Arrow 1933
Onthophagus anatolicus Petrovitz 1962
Onthophagus anchommatus Lea 1923
Onthophagus andalusicus Waltl 1835
Onthophagus anisocerus Erichson 1842
Onthophagus arai Masumoto 1989
Onthophagus asper Macleay 1864
Onthophagus atricapillus d'Orbigny 1908
Onthophagus atrofasciatus D' Orbigny 1905
Onthophagus atrox Harold 1867
Onthophagus australis Guerin-Meneville 1830
Onthophagus azusae Ochi & Kon 2006
Onthophagus babaulti D' Orbigny 1915
Onthophagus beesoni Arrow 1931
Onthophagus beiranus Peringuey 1908
Onthophagus bicolensis Ochi & Kon 2006
Onthophagus bicornis Macleay 1888
Onthophagus bindaree Storey & Weir 1990
Onthophagus binyana Storey & Weir 1990
Onthophagus bonsae Zunino 1976
Onthophagus bornemisszai Matthews 1972
Onthophagus bubalus Harold 1867
Onthophagus bunamin Matthews 1972
Onthophagus bytinskii Balthasar 1960
Onthophagus caesariatus Boucomont 1921
Onthophagus capelliformis Gillet 1925
Onthophagus caprai Frey 1956
Onthophagus carinulatus Harold 1877
Onthophagus cavia Boucomont 1914
Onthophagus chepara Matthews 1972
Onthophagus circulator Reitter 1891
Onthophagus citreum Boucomont 1919
Onthophagus compressus Guerin-Meneville 1855
Onthophagus cyaneoniger D' Orbigny 1902
Onthophagus dellacasai Pittino & Mariani 1981
Onthophagus depilatus D' Orbigny 1913
Onthophagus depressus Harold 1871
Onthophagus diversiformis Boucomont 1914
Onthophagus eschscholtzi Boucomont 1924
Onthophagus falculatus Boucomont 1914
Onthophagus falzonii Goidanich 1926
Onthophagus ferrari Matthews 1972
Onthophagus flavolimbatus Klug 1855
Onthophagus fritschi D' Orbigny 1902
Onthophagus fuscivestis D' Orbigny 1902
Onthophagus gajo Krikken 1977
Onthophagus giraffa Hausmann 1807
Onthophagus granulatus Boheman 1858
Onthophagus gulmarri Matthews 1972
Onthophagus hageni Lansberge 1883
Onthophagus hemipygus Frey 1964
Onthophagus hermonensis Baraud 1982
Onthophagus hildebrandti Harold 1878
Onthophagus hirtus (Illiger 1803)
Onthophagus histeriformis Boucomont 1914
Onthophagus hoplocerus Lea 1923
Onthophagus humpatensis Lansberge 1886
Onthophagus hyalcyon Palestrini & Giacone 1989
Onthophagus impressicollis Boheman 1860
Onthophagus iyengari Arrow 1931
Onthophagus janthinus Harold 1875
Onthophagus jugicola D' Orbigny 1902
Onthophagus juvencus Klug 1835
Onthophagus kanarensis Arrow 1931
Onthophagus kangeanus Paulian 1936
Onthophagus kavirondus D' Orbigny 1915
Onthophagus kirki Frey 1975
Onthophagus kyleensis Frey 1975
Onthophagus lacustris Harold 1877
Onthophagus lamgalio Matthews 1972
Onthophagus laminatus Macleay 1864
Onthophagus latigena d'Orbigni 1897
Onthophagus leanus Goidanich 1926
Onthophagus lemekensis D' Orbigny 1915
Onthophagus lugubris Fahraeus 1857
Onthophagus mamillatus Lea 1923
Onthophagus manya Matthews 1972
Onthophagus massai Baraud 1975
Onthophagus melitaeus (Fabricius 1798)
Onthophagus merdarius Chevrolat 1865
Onthophagus muelleri Novak 1921
Onthophagus mundill Matthews 1972
Onthophagus muticus Macleay 1864
Onthophagus nagpurensis Arrow 1931
Onthophagus nefarius Balthasar 1941
Onthophagus neostenocerus Goidanich 1926
Onthophagus nigellus (Illiger 1803)
Onthophagus nigriventris D' Orbigny 1905
Onthophagus numidicus d'Orbigny 1908
Onthophagus obtusicornis Fahraeus 1857
Onthophagus onorei Zunino & Halffter 1997
Onthophagus opacicollis Reitter 1893
Onthophagus panici Petrovitz 1964
Onthophagus parallelicornis Macleay 1887
Onthophagus parumnotatus Fahraeus 1857
Onthophagus petrovitzianus Pittino 2006
Onthophagus planifrons Frey 1962
Onthophagus praedatus Harold 1862
Onthophagus probus Peringuey 1901
Onthophagus propinquus Macleay 1888
Onthophagus pseudovirens D' Orbigny 1913
Onthophagus pugionatus Fahraeus 1857
Onthophagus punctatus (Illiger 1803)
Onthophagus punthari Storey 1977
Onthophagus quadricuspis D' Orbigny 1908
Onthophagus queenslandicus Blackburn 1903
Onthophagus rhinocerus Gomes Alves 1944
Onthophagus rufosignatus Macleay 1864
Onthophagus rugosicollis Gillet 1925
Onthophagus rugulipennis Fairmaire 1887
Onthophagus semimetallicus Lea 1923
Onthophagus sericatus Reitter 1893
Onthophagus sikkimensis Gillet 1925
Onthophagus spathatus Boucomont 1914
Onthophagus stigmosus D' Orbigny 1902
Onthophagus strabo Reitter 1892
Onthophagus stylocerus Graells 1851
Onthophagus subocelliger Blackburn 1903
Onthophagus suermelii Petrovitz 1963
Onthophagus surdus Boucomont 1925
Onthophagus tamworthi Blackburn 1903
Onthophagus tenebrosus Harold 1871
Onthophagus trapezicornis D' Orbigny 1902
Onthophagus tricolor Boucomont 1914
Onthophagus trigibber Reitter 1893
Onthophagus tweedensis Blackburn 1903
Onthophagus victoriensis Blackburn 1903
Onthophagus vigens Peringuey 1901
Onthophagus vilis Harold 1877
Onthophagus weringerong Storey & Weir 1990
Onthophagus xanthomerus Bates 1887
Onthophagus yaran Storey & Weir 1990
Onthophagus yiryoront Matthews 1972
Onthophagus yungaburra Matthews 1972
Oxysternon macleayi (Nevison 1892)
Oxysternon pteroderum Nevinson 1892
Pachysoma aesculapius (Olivier 1789)
Pachysoma denticollis (Peringuey 1888)
Pachysoma endroedyi (Harrison, Scholtz & Chown 2003)
Pachysoma fitzsimonsi (Ferreira 1953)
Pachysoma gariepinus (Ferreira 1953)
Pachysoma glentoni (Harrison, Scholtz & Chown 2003)
Pachysoma hippocrates (Macleay 1821)
Panelus bakeri Boucomont 1921
Paracanthon trichonotulus Balthasar 1938
Paraphytus africanus Boucomont 1923
Paraphytus foveatus Boucomont 1923
Pedaria biseria Josso & Prevost 2003
Pedaria cambeforti Josso & Prevost 2003
Pedaria conformis Peringuey 1901
Pedaria criberrima Waterhouse 1890
Pedaria decorsei Boucomont 1922
Pedaria dentata Raffray 1877
Pedaria durandi Paulian, Cambefort & Mauchamp 1982
Pedaria estellae Josso & Prevost 2006
Pedaria granulosa Ferreira 1962
Pedaria hanae Balthasar 1963
Pedaria insularis Kolbe 1914
Pedaria juhellegrandi Josso & Prevost 2003
Pedaria morettoi Josso & Prevost 2003
Pedaria ovata Boucomont 1922
Pedaria puncticollis Waterhouse 1890
Pedaria spinithorax Paulian, Cambefort & Mauchamp 1982
Pedaria tenebrosa Balthasar 1937
Pedaria tibialis Josso & Prevost 2006
Pedaridium hirsutum (Harold 1859)
Phanaeus achilles Boheman 1858
Phanaeus bispinus Bates 1868
Phanaeus prasinus Harold 1868
Phanaeus pyrois Bates 1887
Phanaeus sororibispinus Edmonds & Zidek 2012
Platyonitis bicuariensis Ferreira 1976
Proagoderus atriclaviger (d'Orbigny 1905)
Proagoderus biarmatus (d'Orbigny 1908)
Proagoderus bicallosus (Klug 1855)
Proagoderus brucei (Reiche 1849)
Proagoderus extensus (Harold 1878)
Proagoderus plato (Bates 1888)
Proagoderus quadrituber (d'Orbigny 1908)
Proagoderus sapphirinus (Fahraeus 1857)
Proagoderus uluguru (Frey 1975)
Proagoderus versus (d'Orbigny 1913)
Prodontria lewisi Broun 1904
Pseudochironitis stuhlmanni Kolbe 1897
Pseudocotalpa giulianii Hardy 1974
Sarophorus punctatus Frolov & Scholtz 2003
Sauvagesinella palustris Matthews 1974
Scarabaeus aegyptiorum Gerstaecker 1871
Scarabaeus ambiguus Boheman 1857
Scarabaeus asceticus Gillet 1907
Scarabaeus augias (Gillet 1908)
Scarabaeus bornemizzai Zur Strassen 1980
Scarabaeus caffer Boheman 1857
Scarabaeus canaliculatus Fairmaire 1888
Scarabaeus costatus (Wiedemann 1823)
Scarabaeus damarensis Janssens 1940
Scarabaeus deludens Zur Strassen 1961
Scarabaeus difficilis (Zur Strassen 1965)
Scarabaeus ebenus Klug 1855
Scarabaeus gagates (Shipp 1895)
Scarabaeus galenus Westwood 1847
Scarabaeus inoportunus Ferreira 1953
Scarabaeus karrooensis Zur Strassen 1961
Scarabaeus laevistriatus Fairmaire 1893
Scarabaeus namibicus Krajcik 2006
Scarabaeus nigroaeneus Boheman 1857
Scarabaeus ritchiei MacLeay 1821
Scarabaeus rixosus Peringuey 1901
Scarabaeus subaeneus Harold 1869
Scarabaeus xavieri Ferreira 1968
Scatimus onorei Genier & Kohlmann 2003
Scatimus pacificus Genier & Kohlmann 2003
Scatimus simulator Martinez 1988
Scatonomus lauropalui Vulcano & Pereira 1973
Scatonomus viridis Pereira & Martinez 1959
Scatonomus xanthopygus Pereira 1954
Scybalocanthon darlingtoni (Paulian 1939)
Scybalocanthon nigellus (Schmidt 1922)
Scybalocanthon nigriceps (Harold 1868)
Scybalocanthon sexspilotus (Guérin-Méneville 1855)
Scybalocanthon trimaculatus (Schmidt 1922)
Scybalophagus patagonicus Martínez 1953
Scybalophagus plicatipennis (Blanchard 1843)
Scybalophagus rugosus (Blanchard 1846)
Sisyphus caffer Boheman 1857
Sisyphus fasciculatus Boheman 1857
Sisyphus natalensis Balthasar 1968
Sisyphus ocellatus Peringuey 1901
Sulcophanaeus actaeon (Erichson 1847)
Sulcophanaeus auricollis (Harold 1880)
Sulcophanaeus batesi (Harold 1868)
Sulcophanaeus columbi Macleay 1819
Sulcophanaeus imperator (Chevrolat 1844)
Sulcophanaeus miyashitai Arnaud 2002
Sylvicanthon bridarollii Martinez 1949
Sylvicanthon obscurus Schmidt 1920
Temnoplectron bornemisszai Matthews 1974
Temnoplectron boucomonti Paulian 1934
Temnoplectron cooki Reid & Storey 2000
Temnoplectron disruptum Matthews 1974
Temnoplectron finnigani Reid & Storey 2000
Temnoplectron rotundum Westwood 1841
Temnoplectron subvolitans Matthews 1974
Tesserodon feehani Storey 1991
Tesserodon pilicrepus Matthews 1974
Tesserodoniella elguetai
Thyregis relictus Matthews 1976
Thyregis tarsatus Paulian 1933
Tomogonus crassoides Boucomont 1923
Tomogonus crassus (d'Orbigny 1902)
Trichillidium quadridens (Arrow 1932)
Trichillum cordobense
Uroxys angulicollis (Boheman 1858)
Uroxys catharinensis Balthasar 1966
Uroxys coarctatus Harold 1867
Uroxys dilaticollis (Blanchard 1846)
Uroxys elongatus Harold 1868
Uroxys latesulcatus Bates 1891
Uroxys lojanus Arrow 1933
Uroxys monstrosus Balthasar 1940
Uroxys pygmaeus Harold 1883
Uroxys rugatus Boucomont 1928
Uroxys variabilis Robinson 1951
Xinidium davisi Cambefort 1985
Zonocopris machadoi
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wikidata_hierarchy tar gz
Biota
Eukaryota
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Animalia
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Maladera aureola
Maladera bakeri
Maladera businskyorum
Maladera celebensis
Maladera chenzhouana
Maladera crinifrons
Maladera dahongshanica
Maladera davidis
Maladera delicta
Maladera detersa
Maladera eclogaria
Maladera ejimai
Maladera elata
Maladera eluctabilis
Maladera enigma
Maladera eshanensis
Maladera fangchengensis
Maladera fencli
Maladera festina
Maladera floresina
Maladera fulgida
Maladera fuliginosa
Maladera fumosa
Maladera furcillata
Maladera futschauana
Maladera ganglbaueri
Maladera gibbiventris
Maladera graeca
Maladera guangdongana
Maladera guomenshanensis
Maladera guttula
Maladera hajeki
Maladera harmonica
Maladera hauseri
Maladera hsui
Maladera ignava
Maladera immunita
Maladera implicata
Maladera impressithorax
Maladera indica
Maladera inermis
Maladera infuscata
Maladera insanabilis
Maladera insubida
Maladera insularis
Maladera iridicauda
Maladera juntongi
Maladera kamiyai
Maladera kinabaluensis
Maladera krueperi
Maladera kusuii
Maladera lata
Maladera lugubris
Maladera malangeana
Maladera manipurana
Maladera marginella
Maladera mirabilis
Maladera modestula
Maladera moebiusi
Maladera okinawaensis
Maladera okinoerabuana
Maladera opima
Maladera ovatula
Maladera riberai
Maladera robustula
Maladera rubida
Maladera rufopaca
Maladera schoenfeldti
Maladera secreta
Maladera seleuciniensis
Maladera shaowuensis
Maladera shiniushanensis
Maladera singularis
Maladera sinica
Maladera sinistra
Maladera sinuosa
Maladera solida
Maladera sordida
Maladera straminea
Maladera villiersi
Maladera weni
Maladera wipfleri
Maladera wuliangshanensis
Maladera xingkei
Maladera yaeyamana
Maladera yibini
Omaladera
448 additional siblings truncated for brevity.
See the
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wikidata_hierarchy tar gz
Biota
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Outenikwanus
Oxycetonia
Oxysternon
Pachylomera
Pachypodinae
Parabyrsopolis
Paracanthon
Paracryptocanthon
Paragynaecoserica
Paraphacosomoides
Parascarabaeus
Parascatonomus
Parbomonyx
Paroniticellus
Paronitis
Parvuhowdenius
Passalaphodius
Paulosawaya
Peckolus
Pedaridium
Pelidnotopsis
Penalus
Pentanomala
Pereiraidium
Phacosoma
Phaedotrogus
Phaenomeridinae
Phaenomeris
Phanaeus
Philoserica
Pinacotarsus
Plaesiorrhina
Pleuronitis
Plusiotis
Plusiotus
Podoserica
Poecilodiscus
Polylamina
Popillia
Protoleura
Prototroginae
Pseudammoecius
Pseudignambia
Pseudocanthon
Pseudochironitis
Pseudochlorota
Pseudocopris
Pseudocotalpa
Pseudoisonychus
Pseudonthobium
Pseudopedaria
Pseudosaproecius
Pseuduroxys
Pteronyx
Pycnopanelus
Rhipiphoria
Rhyncoserica
Rhyncosymmela
Rutelinae
Saphobiamorpha
Saphobius
Saprosites
Saprosotellus
Saprosties
Sarothermus
Sauvagesinella
Scaptocnemis
Scarabaeinae
Scatonomus
Scatrichus
Sceliages
Scybalocanthon
Scybalophagus
Scythrodes
Sericinae
Shangaia
Simogonious
Sinapisoma
Sisyphus
Sphaerocanthon
Stiptocnemis
Stiptotarsus
Stomanomala
Streblopus
Strehlia
Sulcodrepanus
Sulcophanaeus
Sylvicanthon
Tanzanolus
Taurhina
Temnoplectron
Teraserica
Termitotrox
Tesserodon
Tesserodoniella
Tetraechma
Tetramereia
Theotimius
Thyregis
Tomogonus
Tragiscus
Tribostethes
Trichelasmus
Trichillidium
Trichius
Trichoserica
Trochaloserica
Tropidonitis
Unidentis
Valginae
Vulcanocanthon
Walterantus
Xenocanthon
Xinidium
Xynedria
Yvescambefortius
Zonocopris
Acanthoceridae
Aegialiidae
Alloioscarabaeidae
Aulonocnemidae
Belohinidae
Bolboceratidae
Ceratocanthidae
Cetoniidae
Coprinisphaeridae
Cretoserica
Diphyllostomatidae
Dynastidae
Eremazidae
Euchiridae
Geotrupidae
Glaphyridae
Glaresidae
Hybosoridae
Lithoscarabaeidae
Lucanidae
Ochodaeidae
Orphnidae
Pachypodidae
Pallichnidae
Paralucanidae
Passalidae
Pleocomidae
Rutelidae
Septiventeridae
Trogidae